The Distribution of
Dragonflies And Damselflies (Odonata) in Georgia
Bill Mauffray and Giff Beaton
(Mauffray)
International Odonata Research Institute, % Division of Plant Industry
P.0. Box 147100, Gainesville, FL. 32614< iodonata@bellsouth.net >
(Beaton)
320 Willow Glen Dr, Marietta
GA 30068 giffbeaton@mindspring.com
ISSN 1061-3781
Bulletin Of American
Odonatology: Vol 9, No 2: pp 21-66
10-June 2005
Modified for the web by Bill
Mauffray
Abstract
We present a list of 173 odonate taxa (170 species)
from Georgia. Four taxa are newly added
to the state list: Calopteryx amata,
Argia fumipennis violacea, Enallagma coecum, and
Gomphus australis. Several
species listed in recent publications are removed from the list: Lestes forcipatus, Gomphus crassus,
Gomphus septima, Cordulegaster diastatops, Epitheca spinigera, Erythrodiplax
umbrata, Ladona exusta, and Libellula jesseana. Synonyms and unsupportable older species
records are discussed.
Introduction
A comprehensive distributional study of Georgia
dragonflies and damselflies (Odonata) is presented. Listed are 173 taxa
comprising 170 species, including 51 species (53 taxa) of Zygoptera and 120
taxa representing 119 species of Anisoptera. Data was derived from both
author’s field studies throughout Georgia between 1995 and 2004, examination of
specimen data in several museums, field notes of others, and from literature.
In 1995 a Georgia Dragonfly Survey was organized. A
number of field trips were organized to cover many of Georgia’s counties to
discover new records. Several Dragonfly Society of the Americas (DSA) regional
meetings were held to assist with the survey. The junior author, a resident of
Georgia, criss-crossed the state with both net and camera in hand, accumulating
over 1500 new records during the study period. Meanwhile the senior author
inventoried the collections at the University of Georgia in Athens, Georgia;
the Florida State collection of Arthropods (FSCA) in Gainesville, Florida, and
the National Museum of Natural History (NMNH) in Washington, DC. Data was also
compiled from a number of researchers and an extensive literature review was
performed.
A study of references from surrounding states has
helped to formulate a better understanding of the distribution of Odonata
species within Georgia. References to doubtful records were researched and are
presented and a list of species expected in Georgia is discussed. The extensive
bibliography includes references from Georgia, as well as those from surrounding
states of Alabama, North Carolina, South Carolina, Tennessee, and northern
Florida. The results of this 10-year study have helped fill quite a few
distributional gaps within Georgia. The Georgia list is now comparable with
other eastern states such as Alabama, 173 (Tennessen, et al 1995); Florida, 162
(Dunkle, 1992); New Jersey, 172 (May & Carle, 1996); New York 175
(Donnelly, 1992); Ohio, 156 (Glotzhober, 1995). It is hoped that this list will
stimulate continuing studies within Georgia.
Two web sites (<http://www,afn.org/~iori/galist.htm>
and <http://www.giffbeaton.com/dragonflies.htm>) were established in
conjunction with this survey. Both will continue to be maintained in order to
provide a vehicle for new records to be posted. Also, an annual summary will be
posted in ARGIA, the newsletter of Dragonfly Society of the Americas.
Historical review Click here to see Spreadsheet of Georgia Odonata history (not
part of printed version)
An extensive study of literature records was
performed by the senior author. The historical study of Georgia Odonata has
some interesting turns and twists. Several species which were listed during the
19th century have been either synonymized or removed to doubtful status. The earliest
records found include 4 taxa listed by Burmeister (1839). During the next 50
years, Hagen (1861, 1863, 1874 & 1875), Rambur (1842) and Selys (1850,
1853, 1854a, 1854b, 1858, 1862, 1876, 1878a, 1876b & 1879) added 71 more
taxa and 4 were removed due to synonymy, resulting in 71 taxa on the list by
mid 1890. Many of these early records were based on one or only a few
specimens. Some of these records have
been cited multiple times over the years leading one to believe that there were
many early Georgia records. Several of the doubtful taxa that we have removed
in this report refer to a single record made during this period.
Between mid 1890 and 1923, Bradley
(1914), Calvert (1893 & 1902, 1913), Davis
(1911), Kirby (1890), Martin (1906) Muttkowski (1910, 1915), Ris (1910
& 1911) and Williamson (1923a, 1923b) added 19 taxa and removed one taxon
bringing the total to 88. This cumulative total was increased to 99 by the
addition of 11 species by Root (1924).
By 1954 the total increased to 111 as a result of additions by Byers
(1927b, 1931 & 1937), Montgomery (1947) Needham & Heywood (1929),
Pritchard (1935), Walker (1925 & 1952), Westfall (1943, 1953) and
Williamson (1932, 1934). Six were removed during this period.
In 1955, Needham & Westfall added 11 Anisoptera
and listed a total 77 Anisoptera from Georgia. The cumulative total of Odonata
stood at 122 species. By 1995 the list had grown to 149 with additions from
Bick (1978 & 1990), Carle (1979, 1980, 1982), Donnelly (1973), Dunkle
(1975, 1983), Garrison (1994a), Gloyd (1968), Kormondy (1960) and Louton
(1982). Bennefield (1965), Johnson (1973b) and Westfall and Tennessen (1979)
removed three species from the list.
During the Georgia Dragonfly Survey
period, which began in 1995, thirty-two additional records were published by
Bick (1997), Caldwell (1999), Mauffray (1998, 2003, 2004), Needham, Westfall,
and May (1996) and Westfall and May (2000). The senior author, who assisted
with the distributional data for both of those publications, provided most of
the records in the latter two. One species was removed by Bick (2003). The
total was 180 prior to this publication.
Four new taxa are added here: Calopteryx
amata, Argia fumipennia violacea, Enallagma coecum, Gomphus australis;
and 11 species are removed from the list due to various reasons. With these
adjustments the final list stands at 173 taxa representing 170 species.
Zygoptera total 53 taxa representing the families: Calopterygidae, Lestidae,
Coenagrionidae, and Anisoptera total 80 taxa, including the families
Petaluridae, Aeshnidae, Gomphidae, Cordulegastridae, Corduliidae, and
Libellulidae.
Physiography of Georgia, and Distribution of
Odonata Maps: Physiographic
1 - Physiographic 2 - Counties - Rivers
For several reasons, Georgia is ideally
located to have a large and diverse odonate fauna. In addition to being the
largest state east of the Mississippi (58,910 square miles), its location allows
for a unique mix of northern and southern species. Following the scheme of
Hodler and Schretter (1986), Georgia has five major physiographic regions or
provinces based on a combination of topography, elevation, underlying soils,
and predominant vegetation (Fig. 1). The Blue Ridge Mountains reach into the
northeast corner of the state, and the Cumberland, or Appalachian, Plateau just
brushes the northwest corner. In between the two is the Ridge and Valley
Province. The southern border of all three is the Piedmont region, which covers
the area south to the Fall Line. The line forms the boundary between the
Piedmont and the Coastal Plain. The Coastal Plain covers the remainder of the
state, which is almost 40% of the total area of Georgia.
The Blue Ridge Province, which contains
all of Georgia’s highest mountains and supports the most northern Odonate
fauna, is a group of mountains, ridges, and basins. This is the highest part of
the state, ranging from 480m-1410m (1600’ to 4700’), and is the coldest section
in Georgia. This region also receives the highest rainfall in Georgia. The
habitat consists of high elevation ponds and lakes and associated marshland,
and fast cold streams with medium to high gradients. Not surprisingly, this
region is the extreme southern terminus for many odonate species’ ranges in
eastern North America. One northern species of Zygoptera, Enallagma
hageni, occurs in Georgia only in the Blue Ridge. Lestes eurinus
is almost restricted to this region, although there are several North Carolina
records from the Cumberland Plateau. Archilestes
grandis has been found in Georgia only three times outside this region.
Within the Anisoptera, the
following species are found in Georgia only in the Blue Ridge: Calopteryx amata, Gomphus
(Hylogomphus) adelphus, Gomphus (Gomphus) quadricolor, Lanthus vernalis, Ophiogomphus
edmundo, O. mainensis,
Stylurus scudderi, S. spiniceps, and Somatochlora elongata.
Several of these species are known from only one or two specimens, as would be
expected at the limit of their range. Three other species extend barely west
into the northern Ridge and Valley or Cumberland Plateau, but also reach the
southwestern limit of their range: Boyeria
grafiana and Sympetrum
semicinctum and rubicundulum.
The Cumberland Plateau, in the extreme
northwest corner of the state, is a fairly high and flat plateau dominated in
Georgia by the Lookout and Sand Mountain areas, where elevations range from
240m-550m (800’ to 1800’). This is the remnant of a sandstone plateau, with a
limestone valley between the two. This area isn’t as unique in the south as the
Blue Ridge, but several species of odonates barely reach the state in this
region (or barely into the western edge of the Ridge and Valley). They are Arigomphus villosipes, Dromogomphus spoliatus, Gomphus (Gomphurus) lineatifrons,
and Stylurus notatus (one
historical record).
The Ridge and Valley Province is
situated between these two highland areas. From the edge of the Cumberland
Plateau it extends to the east and south from the Tennessee border south to
about Cartersville, east to about highway US 411, and west to the Alabama
border below the Cumberland Plateau. As the name implies, this is an area of
alternating narrow parallel ridges and wide valleys, and is the least
“mountain-like” area in the extreme northern part of Georgia. It is also the
lowest at 240m-480m (700’ to 1600’), although most of it is on the lower
elevation end of that range. In some ways the Ridge and Valley is similar to
the first two provinces but in many other ways it more closely resembles the
Piedmont to the south. No species are found only in the Ridge and Valley of
Georgia, but there are several species that occur only across these three
northern regions, which comprise about 15% of the land area of the state.
Damselfly species that occur only in these three regions, most of which are at
their most southern distribution, are Calopteryx angustipennis, Chromagrion conditum (also a few
records in northern Piedmont), and Enallagma
aspersum. Anisopterans found only in this northern section are Gomphus (Gomphurus) consanguis, Cordulegaster erronea, and Helocordulia uhleri. Although their
ranges are poorly known, three species of River Cruisers may belong to this
group in Georgia: Macromia
alleghaniensis, M. illinoiensis
illinoiensis, and M. margarita.
The Piedmont Region is the next region
to the south, and stretches from the Alabama border in the west to the South
Carolina border in the east and south to the Fall Line. The term Piedmont comes
from an Italian word meaning “Foot of the Mountain,” which describes the
northern edge of the Piedmont. The Fall Line is named after the line where
rivers passing from the higher elevation of the Piedmont to the lower, flatter
Coastal Plain have waterfalls and rapids. This line marks the southern boundary
of the Piedmont, and numerous major cities are built along it, including
Columbus, Macon, Milledgeville, and Augusta. The Piedmont has some northern
features but some southern features also; it is characterized by rolling
topography and larger rivers than those in areas to the north, plus a few
isolated monadnocks (isolated mountain remnants). Elevations are mostly lower,
ranging from 150m (500’) at the Fall Line to 967m (3173’) at Mount Yonah, the
tallest monadnock. The average elevation on the northern edge of the Piedmont
is 360m (1200’). Temperatures are intermediate for Georgia. A number of species
reach their southernmost distribution in the Piedmont, the most notable of
which are Aeshna umbrosa, Gomphus (Gomphurus) rogersi, Gomphus (Hylogomphus) parvidens, Ophiogomphus incurvatus, Stylogomphus albistylus, Stylurus laurae, and Somatochlora tenebrosa. The latter
two species have been found south of the Piedmont in nearby states, and we
expect to find a few in south Georgia as well. Many species also reach their
most northern or western limit in the Piedmont, which in many cases is the
farthest inland they reach anywhere in their range. These include Gomphaeschna antilope, Aphylla williamsoni and Orthemis ferruginea (the latter two
species extending their ranges into the southern Piedmont in recent years).
Along much of the Fall Line is a series
of deep sand ridges forming the Fall Line Sandhills. This area is the border
between the lower elevation, marine sediments making up the Coastal Plain and
the rockier, older Piedmont; it was the ocean shoreline during the Mesozoic
Era. One species found only in the Sandhills so far is Enallagma davisi; we do not expect it to be found anywhere else
in the state. A couple of other species may be mostly restricted to this region
also: Gomphus (Gomphus) diminutus
(only one record near Augusta, at the westernmost limit of its range) and Nannothemis bella. Also in this
restricted habitat (only in Taylor County so far) is a puzzling group of Gomphus (Hylogomphus) geminatus
records, well away from its known range on the Gulf Coast. Further study is
planned to search for records of this species in between these two areas.
South of the Fall Line lies the Coastal
Plain Region, bordered by Alabama on the west, Florida to the south, and the
Atlantic Ocean or the Savannah River next to South Carolina on the eastern border.
The Coastal Plain is fairly flat and warm, ranging from sea level to 150m
(500’), and has larger rivers with attendant wide areas of floodplain. Several
clubtails are found most commonly along these rivers, such as Gomphus (Gomphurus) dilatatus and hybridus. There are actually two
slightly different coastal plains in Georgia: the Atlantic Coastal Plain
includes all river basins emptying into the Atlantic, and the Gulf Coastal
Plain includes all river basins emptying into the Gulf of Mexico (mostly southwestern
Georgia). The Okefenokee divides the two, with rivers flowing from it in both
directions. Most of the species of interest in this region are those that reach
their northernmost points here, and many of these have a mostly Florida range.
Some barely reach Georgia at all, such as Enallagma coecum, E. pollutum, and Gynacantha nervosa.
Other notable odonate examples include Enallagma
pallidum, Triacanthagyna trifida,
Arigomphus pallidus, Gomphus australis, Epitheca semiaquea, E. sepia,
E. stella, Celithemis amanda and Miathyria marcella. Two species are
found either immediately along the coast (Erythrodiplax berenice) or along the coast and slightly inland
(Libellula needhami).
Acknowledgements
George and the late Juanda Bick inventoried the FSCA
collection and provided personal data, moral support, and the foundation and
motivation to undergo this study. Nancy Adams assisted in inventorying the NMNH
collection. Cecil Smith provided access to the University of Georgia collection
in Athens Ga. The late Dr. Minter J. Westfall Jr. provided additional moral
support as well as specimens from several counties. Additional specimen, dates
and photographic records were provided by many others during the study
including but not limited to: Peter Allen, Robert Behrstock, Thomas Donnelly,
Sid Dunkle, Jerrell Daigle, Sandy Garrett, Alan Harvey, Steve and Mary Jane
Krotzer, Steve Parrish, Dennis Paulson, David Small, Dirk Stevenson, Ken
Tennessen, Linden Trial, Michael Veit, Minter J. Westfall Jr., and especially Marion
Dobbs; plus excellent photographic data from James Flynn, Francis Michael
Stiteler, Rusty Trump and others who attended the DSA gatherings. Thanks to Ken
Tennessen for reviewing the final draft and making some valuable suggestions.
Special recognition to Esther Mauffray, the senior author’s wife who was very
patient during the final writing of this study, and to Becky Beaton, wife of
the junior author and highly tolerant and supportive during many years of
studying Georgia Odonates.
The families are listed after Garrison
(1997-2004) and all matters of nomenclature are based on his 2004 revision. We
differ only by not differentiating the two subspecies of Epitheca
(Epicordulia) princeps. The species are listed in alphabetical
order followed by the common name adopted by the Dragonfly Society of the
Americas (DSA). The first record is
marked with an “*.” Previous “state only” records are listed first in
chronological order. These include literature references that list only Georgia
with no county or locality records. If
Georgia is included in a range description (i.e. Florida to Maryland) the
reference is not cited. If Georgia is named in the range description (ie,
Georgia to Maryland) then the record is included. If a range map covered a
portion of Georgia, the reference is not included; however if the map is a dot
map, such as Louton (1982) or Donnelly (2004a, 2004b & 2004c), and shows
explicit Georgia records, then it is considered a valid reference for the
state. To save space the following citations are abbreviated: Georgia Water
Quality Control Board (1971) is GWQ (1971), Needham and Westfall (1955) is
N&W (1955); Needham, Westfall & May (2000) is NW&M (2000) and
Westfall and May (1996) is W&M (1996).
County records follow and are listed alphabetically,
each with codes (see legend below) and/or literature references. A county map
is provided (Fig. 2). Records from multiple sources are listed. In some cases a
multiple-record reference may refer to the same specimen which was cited in the
literature and then again in a particular collection. The authors collected
many of the specimens upon which the FSCA, IORI, and NMNH records are
based. Multiple references better
support the validity of records, especially if one or more of the records was a
sight record. A brief note about the species habitat with regards to Georgia is
included followed by the earliest and latest recorded or known dates for adults
enclosed in brackets.
Legend: codes within parentheses ( ).
"*" = first
published record; pers. comm.= personal
communication
AH1= Alan Harvey, pers.
comm. (GDS 2001 notes)
AH2= Alan Harvey, pers.
comm. (GDS 2002 notes)
AH3= Alan Harvey, pers.
comm. (GDS 2003 notes)
BB= Bob Barber, pers. comm.
DB42= Borror, 1942
Br14= Bradley, 1914
Bu39= Burmeister, 1839
By27c= Byers, 1927c
By31= Byers, 1931
By39= Byers, 1939
Ca04= Calvert, 1904
Ca12= Calvert, 1912
Ca13= Calvert, 1913
Ca79= Carle, 1979
Ca80= Carle, 1980
Ca82= Carle, 1982
Ca83= Carle, 1983
Ca98= Calvert, 1898
Ca99= Caldwell, 1999
CC89c= Cook, 1989c
CJ72= Johnson, 1972
CJ73a= Johnson, 1973a
CJ73b= Johnson, 1973b
CJ74= Johnson, 1974
Cr55b= Cross, 1955b
Da11= Davis, 1911
DC89b= Cuyler, 1989b
DP= Dennis Paulson, pers.
comm.
DG= David Gottleib, pers.
Comm.
DG40= Gloyd, 1940
DG68= Gloyd, 1968
DS= Dirk Stevenson, pers.
comm.
DS2= Dirk Stevenson, 2002 records
DT94= Donnelly & Tennessen ,1994
ET= Eran Tomer, photo 2004
FS2= Francis Michael Stiteler, photos with data 2002
FSCA= FSCA/IORI collections, Gainesville, FL
GB= Giff Beaton, photos and data 2000 and earlier
GB1= Giff Beaton, photos and data from 2001
GB2= Giff Beaton, photos and data from 2002
GB3= Giff Beaton, photos and data from 2003
GB4= Giff Beaton, photos and data from 2004
GB5= Giff Beaton, photos and data from 2005
GBi= Bick, pers. comm.
GBi03= Bick, 2003
GBi78= Bick, 1978
GBi83b= Bick, 1983b
GBi90= Bick, 1990
GBi97= Bick, 1997
GDS3 =Gerogia Dragonfly survey SE DSA Meeting, 22-24 May, 2003.
GWQ71= Georgia Water Quality Board 1971
Ha61= Hagen, 1861
Ha63= Hagen, 1863
Ha74= Hagen, 1874
Ha75= Hagen, 1875
JB3= Jeff Biller, pers.
comm. 2003
JD= Jerrell Daigle, pers.
comm.
JD92b= Daigle, 1992b
JF= James Flynn, photos with data 2002 and earlier
JF3= James Flynn, photos and data from 2003
Ko60= Kormondy, 1960
KP87= Kondratieff & Pyott, 1987
KT= Ken Tennessen, pers.
comm.
KT77= Tennessen, 1977
KT98= Tennessen, 1998
Lo82= Louton, 1982
LT= Linden Trial, pers.
comm. 2004
MD3= Marion Dobbs, photos and data from 2003
MD4= Marion Dobbs, photos and data from 2004
Mu15= Muttkowski, 1915
MV3= Michael Veit, pers.
comm. 2003
MW41= Westfall, 1941
MW43= Westfall, 1943
MW9= Westfall, Coll in 1997-1998, specimens in FSCA
OO= Odes of Okefenokee (web site); Dave Small, 2003
PA= Peter Allen, pers.
comm., 1993-95 records
Pr35= Prichard, 1935
RB3= Robert Behrstock, pers.
comm. 2003
RB= Robert Behrstock, pers.
comm. 1998
RG94a= Garrison, 1994a
RG94b= aGarrison, 1994b
Ro24= Root, 1924
RT= Rusty Trump, photos from 2002
RW67= Roback & Westfall, 1967
SD= Sid Dunkle, pers. comm.
SD00= Dunkle, 2000
SD75= Dunkle, 1975
SD83= Dunkle, 1983
Se53= Selys, 1853
Se62= Selys, 1862
Se76= Selys, 1876
SG3= Sandy Garrett, pers.
comm. 2003
Sh30= Shortess, 1930
SK= Steve & Mary Jane Krotzer, pers. comm..
SK3= Steve Krotzer, pers.
comm. 2003
SK4 =Steve Krotzer, pers.
comm. 2004
SP= Steve Parrish, pers.
comm. 2004
SR= Steven Roble, pers.
comm.
Ta67= Tai, 1967
TD= Thomas Donnelly, pers.
comm.
TD73= Donnelly, 1973
UG= University of Georgia Collection, Athens, GA.
UN= US Museum of Natural History, Washington , DC.
Wa25= Walker, 1925
Wa52= Walker, 1952
Wi14= Williamson, 1914
Wi23a= Williamson,1923a
Wi23b= Williamson, 1923b
Wi32= Williamson, 1932
Wi34= Williamson, 1934
WM03= Mauffray, 2003
WM04= Mauffray, 2004
WM95= Westfall & Mauffray, 1995
WM95a= Mauffray, 1995a
WM98= Mauffray, 1998
WMC= Mauffray collection (many now in FSCA, IORI, LSU, and NMNH)
ZYGOPTERA
Calopteryx amata (Hagen,
1889). Superb Jewelwing. State: Donnelly (2004c)*.
Union (SD).
The Union County record is:
Toccoa River at Forest Road 69, female, 22 Aug 1985, Sid Dunkle. [Only date
recorded: Aug 22].
Calopteryx angustipennis (Selys, 1853). Appalachian Jewelwing. State: Selys (1853*, 1854a, 1859, 1879a) also as elegans,
Hagen (1861, 1863, also as C. splendens, 1874, 1875, 1889), Kirby (1890)
as Agrion elegans, Banks (1892), Montgomery (1947), Johnson (1974), Muttkowski
(1910), N&H (1929), W&M (1996), Tennessen
(1998), Donnelly (2004c).
Dade
(GDS3), Gilmer? (Se53* type), Murray (UG, UN, KT, SK, KT98, WMC, GB4), Walker
(GDS3).
All records prior to 1996 refer to the single Selys
(1853) type specimen record. Johnson
(1974) discussed
the locality data from Selys’ specimen (collected by John Abbot in the 1700s).
Johnson concluded the specimen was from either Burke or Screven County, since
there is a “Brier” Creek running through those counties. The Delorme Georgia
Atlas & Gazetteer (1998) lists a “Brier Creek” which begins near Thompson,
Georgia and runs southeasterly through Burke and Screven Counties into the
Savannah River about 30 miles upstream from Savannah, Georgia. Williamson
(1932) describes Brier Creek in detail (see Stylurus laurae); this area
contains unlikely habitat for this species.
Johnson (1974) says its range “correlates largely with mountain or
up-land type streams.” Hagen (1861) listed it from “Brier Creek” but Hagen
(1863, 1874, 1875, 1889) listed it from “Briar Creek.” To further confuse the
issue, Selys (1879a) lists it from “Brien” Creek. Delorme (1998) also lists
“Briar Creek” in Gilmer County south of Cherry Log off of US 76. It is more
likely that the specimen came from this area. It is uncommon in cold streams in far north Georgia [Apr 18 - Jun 26].
Calopteryx dimidiata Burmeister, 1839. Sparking Jewelwing. State: Selys (1854a*), Hagen (1861, 1863, 1874, 1875,
1889), Banks (1892), Calvert (1906), W&M (1996), Donnelly (2004c).
Bartow
(CJ73b, UG), Berrien (GB3), Bibb (CJ73b), Brantley (CJ73b), Burke (By31, CJ73b,
UN, Wi34 as apicale), Charlton (Br14 as apicale), Chatham (GB),
Clarke (UG), Clinch (WMC), Coffee (CJ73b), Colquitt (CJ73b), Crawford (UG, UN),
Decatur (CJ73b), Dooly (By31), Echols (WMC), Effingham (UG), Elbert (GB), Evans (DP), Floyd (MD4), Gordon
(UN), Gwinnett (By31, CJ73b), Haralson (MD4), Harris (UG), Houston (UG), Jefferson
(CJ73b, Wi34 as apicale), Lee (CJ73b), Lowndes (CJ73b), Morgan (UG, UN),
Peach (UG), Pickens (UG), Pierce (CJ73b), Rabun (MD4), Randolph (GB2), Schley
(GB3, UG), Sumter (UG), Taliaferro (GB), Tattnall (DS), Taylor (JD, RB3, UG),
Telfair (DP), Tift (GB3), Walton (UG), Ware (CJ73b), Wayne (By31, CJ73b),
Whitfield (UG), Wilkinson (DP).
Fairly common in sandy
streams; uncommon in small rivers statewide. It is more common below the fall
line but rare in the northern parts of the state. [April 3 - Sep 22].
Calopteryx maculata (Beauvois, 1805). Ebony Jewelwing.
State: Selys (1854a*), Hagen (1863, 1874, 1875, 1889), W&M (1996), Donnelly
(2004c).
Atkinson
(DP), Bartow (GB, UG, WMC), Burke (Wi34), Butts (MD4), Calhoun (MV3), Catoosa
(GB2), Charlton (Br14, OO), Chattooga (GB, UG), Cherokee (GB2), Clarke (CJ74,
UG), Cobb (GB), Coffee (DP), Columbia (UN), Dade (GB2), Decatur (UG), Dooly
(By31, CJ74), Echols (WMC), Elbert (GB), Fannin (GB), Floyd (UG), Forsyth (UN),
Franklin (WMC), Fulton (HS4), Gilmer (MD4), Gordon (UG, UN), Habersham (GBi),
Haralson (GB2), Harris (GB1, UG), Heard (GB2), Houston (UG), Jackson (UG),
Jasper (GB2), Jefferson (By31, CJ74, Wi34), Jones (WMC), Lamar (UG), Lee (CJ74,
Ro24), Lowndes (UG), McDuffie (Wi34), Meriwether (MD4), Mitchell (JF4), Murray
(GB, KT, UN, WMC), Newton (LT), Oconee (MD3), Oglethorpe (UG), Paulding (GB),
Pickens (GB), Pike (MD4), Polk (UG, UN), Rabun (CJ74), Richmond (UN), Rockdale
(GB), Sumter (UG, UN), Talbot (GB2), Taliaferro (GB), Taylor (JD, RB3), Tift
(UG), Towns (UG), Union (UG), Upson (GB1, GB2), Walker (GDS3), Wayne (By31,
CJ74), Webster (UG), White (KT, Wi34), Whitfield (Ha61, UN), Wilkes (Wi34),
Wilkinson (DP).
Common statewide, mostly in
small streams but can be found in almost any habitat. [Apr 8 - Oct 22].
Hetaerina americana (Fabricius, 1798). American Rubyspot. State: W&M (1996), Donnelly (2004c).
Bartow
(CJ73a*, UG), Carroll (GB), Catoosa (GB), Clarke (UG), Cobb (GB2), Coweta (GB),
Dade (GB2), Floyd (GB2), Fulton (HS4), Gordon (MD4), Laurens (GB), Madison
(CJ73a*), Monroe (CJ73a*), Oconee (UG), Paulding (MD4), Pike (CJ73a, UG), Polk
(GB2), Rockdale (CJ73a*, UG), Talbot (GB2), Towns (GB4), Upson (GB1, GB2),
Walton (CJ73a*, UG).
Uncommon in streams above the
fall line. [May 19 - Oct 2].
Hetaerina titia (Drury, 1773). Smoky Rubyspot. State: Selys (1853*, 1854a), Hagen (1861,
1863, 1874, 1875) and Banks (1892) as H. septentrionalis and H. tricolor, Kirby (1890) as H.
septentrionalis, Calvert (1893, 1906) as H. tricolor, Muttkowski
(1910) also as H. limbata and H. septentrionalis, W&M (1996),
Donnelly (2004c).
Bibb (CJ73a), Brantley (CJ73a), Burke (CJ73a, UN, Wi34), Charlton
(Br14, also as tricolor), Clinch (UG), Coweta (GB2), Decatur (CJ73a,
UG), Early (GB4), Emanuel (MD4), Floyd (By31, CJ73a), Gilmer (GB3), Gordon
(MD4), Haralson (GB2), Heard (GB2), Houston (GB3), Jeff Davis (MD4), Laurens
(GB3), Lee (GB2), Long (GB4), McDuffie (CJ73a, Wi34), Montgomery (MD4),
Paulding (MD4), Pierce (CJ73a), Schley (GB4), Tattnall (GB1), Telfair
(MD3),Twiggs (GB3), Union (CJ73a, Wi34).
Uncommon
throughout the state, in moving water, medium-sized streams to large rivers. [May 14 - Nov 4].
Archilestes
grandis (Rambur, 1842) Great Spreadwing. State: Donnelly (2004c).
Clarke (UG), DeKalb (ET),
Fulton (DG, HS4), Hall (Ca99*), Rabun (GB3), White (Dana Denson per. comm.).
Rare,
recorded only in the northeast part of the state, near streams. [Aug 10 - Nov 3].
Lestes australis Walker,1952. Southern
Spreadwing. Previously known
as L. disjunctus australis (Donnelly, 2003). State: W&M (1996),
Donnelly (2004c).
Baldwin (UG), Bibb (UG), Bulloch (DP), Camden (UN), Clarke (UG),
Clinch (GB3), Cook (WA52*), Dade (GDS3), Dooly (GB2), Dougherty (UG), Echols
(GB3), Evans (WMC, KT), Floyd (MD4), Glynn (WMC), Houston (WA52*), Lamar (UG),
Laurens (GB2, GB3), Long (GB3, WMC), Murray (WMC), Paulding (GB, GB1), Tattnall
(WMC), Telfair (WMC), Tift (UG, WA52*), Toombs (KT), Walker (GDS3), Wheeler
(DP).
Common at ponds,
lakes and marshes throughout the state. [Apr 1 - Nov 13].
Lestes eurinus
Say, 1839. Amber-winged Spreadwing. State: Donnelly (2004c).
Dade (WM04*).
So far known from only the one pond record in extreme northwest
Georgia: Trenton, Cloudland Canyon State Park, pond, 34°49.58’N 85°28.61’WMC, 23 May 2003, 1 male. Coll. G.
Beaton [Only date recorded: May 23].
Lestes inaequalis Walsh, 1862. Elegant Spreadwing. State: W&M (1996*), Donnelly (2004c).
Clarke
(UG), Early (MV3), Habersham (J), Morgan (MV3, DP), Murray (GB3), Richmond
(UN), Wayne (DP), Wilcox (GB2).
Uncommon to rare, scattered
throughout the state in marshes [Apr 11 -
Sep 2].
Lestes rectangularis Say,1839. Slender Spreadwing. State: Banks (1892), Calvert (1893), Montgomery
(1948), W&M
(1996), Donnelly (2004c).
Catoosa
(H), Chatham (Ha61*, Ha63, Se62), Dade (GDS3), Floyd (MD4), Gilmer (GB3), Lee
(Ro24), McDuffie (Wi34), Paulding (GB, GB1), Tattnall (UN) Toombs (UN), Towns
(KT), Wayne (DP), White (SD83), Whitfield (Ha74).
Uncommon to rare, scattered
throughout the state in ponds, marshes, and slow streams. [May 6 - Sep 28].
Lestes vidua Hagen, 1861. Carolina
Spreadwing. State: W&M (1996*),
Donnelly (2004c).
Bibb (UG), Early (GB3), Gwinnett (GB2), Laurens (GB2), Wheeler
(FSCA).
Rare throughout
state at ponds except absent from north Georgia, perhaps overlooked. [Mar 14 - Oct 4].
Lestes vigilax
Hagen in Selys, 1862. Swamp Spreadwing. State: W&M (1996), Donnelly (2004c).
Atkinson (MD4, DP), Bartow (MD4), Bulloch (DP),
Burke (UN, Wi34), Carroll (GB3), Catoosa (GB2), Charlton (UG, PA), Chattooga
(MD4), Colquitt (MD4), Dade (GDS3, GB2), DeKalb (UG), Emanuel (GB4), Floyd
(MD4), Gordon (MD4), Gwinnett (PA), Habersham (GBi), Jones (WMC), Laurens (GB1,
GB4, PA, WMC), Lee (Ro24*), Long (DP), Lumpkin (WMC), McDuffie (UN, Wi34),
Morgan (MD4), Murray (WMC), Oglethorpe (UG), Rabun (GB3), Sumter (GB2, RB),
Taliaferro (GB2), Taylor (GB3), Thomas (UN), Towns (KT), Treutlen (MD4), Walker
(GDS3), Ware (RB, UG), Wayne (DP).
Fairly common throughout the
state at marshes, ponds, and lakes. [Apr 4 - Nov 15]
Coenagrionidae
Amphiagrion saucium
(Burmeister, 1839). Eastern Red Damsel. State: W&M (1996*), Donnelly (2004c).
Bartow (GB1), Burke (GB3), Lumpkin (UG),
Meriwether (UG), Murray (SK), Richmond
(FSCA), Talbot (JD).
Uncommon in very small streams
and seeps above the fall line. [Apr 13 - May 13]
Argia apicalis
(Say, 1839). Blue-fronted Dancer. State: W&M (1996), Donnelly (2004c).
Appling (CJ72), Baldwin (UN), Bartow (GB1, UN), Ben
Hill (GB1), Bibb (UG), Brantley (CJ72), Brooks (GB1), Burke (By31), Carroll
(GB, UN, WMC), Catoosa (GB), Charlton (UG), Clarke (UG), Cook (GB1), Coweta
(GB), Crisp (GB2), Dade (GDS3), Decatur (UG, UN), Dooly (By31), Dougherty
(RW67), Douglas (GB), Early (GB3, SK4), Emanuel (GB1), Fannin (UN), Fayette
(KT), Floyd (GB, GB1, By31), Fulton (UG), Gilmer (GB3), Gordon (GB2), Grady
(GB1), Greene (MD4), Habersham (GBi, GB1, UN), Harris (GB), Heard (GB2), Jeff
Davis (MD4), Jefferson (UN), Jones (GBi), Lamar (GB), Laurens (By31, GB1, UN),
Lee (CJ72, Ro24), Lowndes (CJ72), McDuffie (Wi34), Meriwether (GB1), Miller
(GB), Mitchell (CJ72, UN), Monroe (MD3), Montgomery (GB1, GB2), Morgan (MV3),
Paulding (GB2), Pickens (MD4), Pierce (WMC), Polk (GB, GB2), Putnam (CJ72, UG,
WMC), Rabun (GBi), Richmond (UN), Screven (CJ72), Stephens (GBi, GB2, UN),
Sumter (RB), Tattnall (UN), Telfair (DP), Toombs (UN), Treutlen (GB1), Twiggs
(GB3), Washington (MD4), Wheeler (GB1), White (GB1), Whitfield (UG).
Common across the state at
rivers and large streams, rarely at lakes. [Apr 21 - Oct 2]
Argia bipunctulata
(Hagen, 1861). Seepage Dancer. State: Hagen (1861 type*, 1863, 1874), Selys (1865), Banks (1892),
Calvert (1893), W&M (1996), Donnelly (2004c).
Burke (GB3, Wi34), Cobb (GB), Crawford (UG),
Habersham (GBi, UN), Jefferson (By31), Lee (Ro24), Paulding (GB2), Richmond
(UG), Taylor (GB3).
Uncommon, local; scattered
across the state at seeps, marshy edges and wet grassy areas [May 16 - Sept 26]
Argia fumipennis atra Gloyd, 1968. Black Dancer. State: Donnelly (2004c).
Charlton (DG68*).
Rare in extreme southeast
Georgia at lakes, ponds and rivers. [No date information].
Argia f. fumipennis (Burmeister, 1839).
Smoky-winged Dancer.
State: Hagen (1863*, 1874), Selys (1865), Banks (1892), Calvert (1902), Muttkowski (1910), N&H (1929), Gloyd (1968),
Donnelly (2004c).
Appling (UG), Baker (UN), Bibb (UG), Bryan (UN),
Burke (GB2, UN, Wi34), Carroll (GB, WMC), Charlton (UG), Chattahoochee (DP),
Chattooga (GDS3), Cherokee (GB2), Clarke (UG), Cobb (GB), Coffee (DP), Coweta
(GB2, UG), Crawford (GB, UG, UN), Dade (GDS3), DeKalb (UG, UN), Dooly (By31),
Early (GB2, UG), Elbert (GB), Fayette (KT), Franklin (WMC), Fulton (HS4),
Gordon (GB2), Grady (UN), Gwinnett (By31, GB), Habersham (GBi, UN), Haralson
(WMC), Harris (GB1), Houston (UG, UN), Jasper (GB2), Jefferson (UG, Wi34),
Jenkins (UG), Jones (WMC), Laurens (GB1), Lee (Ro24), Lowndes (UG), Marion
(JF3), McDuffie (UN, Wi34), Meriwether (GB1, GB2), Monroe (GBi), Morgan (UG),
Oglethorpe (UG), Paulding (GB2), Peach (UG), Pickens (GB), Polk (GB2), Pulaski
(UN), Rabun (GB3), Richmond (GBi, UG, UN ), Rockdale (GB), Schley (GB3),
Stephens (GB2), Sumter (MD4, RB), Taliaferro (GB2), Tattnall (UG, WMC), Taylor
(GB3, RB3, SG3, UG, UN), Telfair (DP, UG), Thomas (UG), Tift (GBi), Troup (GB),
Walker (GDS3), Washington (UG), Wayne (DP), Wilkinson (DP).
Common below the middle
Piedmont at lakes, ponds and rivers. Gloyd
(1968) found 1 from Charlton intermediate with A. f. atra. [Apr 11 - Oct 27]
Argia f. violacea
(Hagen, 1861)* Violet Dancer. State: Donnelly (2004c).
Bartow (UG, UN ), Catoosa (MD4), Chattooga (GDS3),
Clarke (UG), Dawson (GB4, MMW9), Fannin (UN), Floyd (MD4), Gilmer (GB3), Gordon
(UN), Habersham (GB1), Hall (FSCA, MMW9), Murray (JD, UN), Paulding (GB1), Polk
(GB2), Union (GB1, GB3), Walker (GDS3), White (GB1), Whitfield (GB2, UG).
All published records list this as a species; we
follow Gloyd’s (1968) assignment as a subspecies. It is common from the middle
Piedmont north, at lakes, ponds and rivers. Intermediates with A. f.
fumipennis are uncommon throughout north Georgia. [May 17 - Oct
8]
Argia moesta
(Hagen, 1861). Powdered Dancer. State: W&M (1996), Donnelly (2004c).
Appling (GB2), Bartow (GB1, MD4), Brantley (DP),
Brooks (GB1), Burke (GB2, Wi34), Butts (WMC), Carroll (GB), Catoosa (GB),
Chattahoochee (UG), Cherokee (UG), Clinch (WMC), Cobb (GB2), Dade (GB2),
Decatur (UG, UN), DeKalb (UN), Dooly (By31), Dougherty (RW67), Early (SK4,
MD4), Echols (GB4, MD4), Effingham (GB2), Floyd (GB1), Gilmer (GB), Gordon
(GB2, UG), Grady (GB1, UG), Gwinnett (By31), Habersham (UN), Haralson (GB2,
MD4), Harris (UG), Heard (GB2), Houston (UN), Jeff Davis (MD4), Jefferson
(GB2), Lee (GB2, Ro24*), Long (GB4), Lowndes (MD4), Macon (GB2), McDuffie
(Wi34), Mitchell (GB2), Monroe (MD3), Montgomery (GB1, GB2), Murray (UN),
Paulding (GB1), Peach (UG), Pierce (JF4),
Polk (GB2),
Quitman (GB2), Randolph (GB2), Richmond (UN), Screven (DP), Sumter (GB2),
Talbot / Upson (GB2), Tattnall (DS, GB4), Telfair (DP), Terrell (UG), Twiggs
(GB3), Upson (GB1), Walker (GDS3), Walton (UG), Ware (RB), Wayne (DP), Wheeler
(GB1), Whitfield (MD3), Wilcox (GB2).
Common throughout the state at
rivers and large streams, uncommon at smaller streams. [May 11 - Nov 3]
Argia sedula
(Hagen, 1861). Blue-ringed Dancer. State: W&M (1996), Donnelly (2004c).
Bartow (GB2),
Berrien (GB3), Butts (WMC), Catoosa (GB), Cherokee (GB2), Clarke (UG),
Coffee (GB3), Coweta (UG), Dade (GB2), Decatur (UG, UN), DeKalb (UN), Early
(GB2, GB3), Fayette (KT), Floyd (GB2), Gordon (GB2), Habersham (UN), Haralson
(GB2), Heard (GB2), Jackson (UN), Jeff Davis (MD4), Jones (GBi), Lee (GB2,
Ro24*), Long (GB4), McDuffie (Wi34), Monroe (GB3, MD3), Montgomery (GB1),
Morgan (DP), Paulding (GB2), Polk (GB2), Screven (DP), Sumter (RB), Talbot /
Upson (GB2), Tattnall (GB4), Telfair (GB3, MD3), Tift (GB3), Walker (GDS3),
Wheeler (GB1), Whitfield (MD3).
Common at rivers and streams
throughout the state except uncommon in the eastern coastal plain, and rare at
ponds. [Apr 18
- Oct 11]
Argia tibialis
(Rambur, 1842). Blue-tipped Dancer. State: Hagen (1863*) as A. fontium, (1874), Selys (1865),
(Banks (1892), W&M (1996), Donnelly (2004c).
Atkinson (WMC), Baker (MV3), Bartow (GB1, UN),
Berrien (GB3), Bibb (UG), Brantley (BB, DP), Brooks (GB1), Burke (By31, Wi34),
Butts (WMC), Calhoun (UG), Catoosa (GB2), Charlton (DP), Chatham (GB), Cherokee
(GB2), Clarke (UG), Clay (GB2), Clayton (UG), Clinch (TD, WMC), Cobb (GB, GB1),
Coffee (DP), Columbia (UN), Cook (GB1), Coweta (GB2), Crawford (UG), Dade
(GDS3, GB2), Decatur (UG, UN ), DeKalb (UG), Dooly (By31), Dougherty (GB2),
Early (GB3, SK4), Echols (WMC), Effingham (GB2), Emanuel (GB1, GB4), Floyd
(By31), Franklin (UG, WMC), Fulton (UG), Gilmer (GB2), Grady (UN), Haralson
(GB2), Harris (UG), Houston (UG, UN), Jackson (UN), Jefferson (By31, GB2, UG,
Wi34), Jones (WMC), Lanier (UG, UN), Laurens (By31, UN, WMC), Lee (Ro24),
Lowndes (UG, WMC), Macon (GB2), McDuffie (Wi34), Mitchell (GB2), Morgan (MV3),
Newton (UG), Paulding (GB2), Pulaski (UN, WMC), Quitman (GB2), Schley (RB3, UG,
UN), Screven (DP), Sumter (RB, UG, UN), Taylor (GB3, RB3, UG, UN), Telfair
(WMC), Terrell (UG), Thomas (UG, UN), Tift (GB3), Treutlen (GB1), Walker
(GDS3), Wayne (DP), Whitfield (MD3, UG), Worth (UN).
Fairly common across the
state, mostly at rivers and streams. [Mar 26 - Oct 13]
Argia translata
Hagen in Selys, 1865. Dusky Dancer. State: W&M (1996), Donnelly (2004c).
Catoosa (GB), Cobb (GB2), Dade (GB2), Floyd (GB2),
Habersham (UN), Haralson (GB2), Heard (GB2), Madison (RG94a*), Meriwether
(MD4), Murray (GB2), Pickens (MD4, UG), Talbot / Upson (GB2), Whitfield (UG).
Fairly common on streams and
rivers above the fall line; rare south of there, and more common in the
northwestern part of the state. [Jun 13 - Sep 19]
Chromagrion conditum
(Selys, 1876). Aurora Damsel. State: W&M (1996*), Donnelly (2004c).
Bartow (GB1), Clarke (UG), Crawford (UG), DeKalb
(UG), Fannin (UG), Gilmer (WMC), Murray (KT, SK), Rabun (FSCA), Talbot (JD),
Walker (GDS3), Whitfield (UG).
Uncommon above the fall line
but fairly common in the upper Piedmont and mountains in the marshy edges of
ponds and small streams. [May 2 - Jun 18]
Enallagma aspersum
(Hagen, 1861). Azure Bluet. State: W&M (1996*), Donnelly (2004c).
Catoosa (UN), Dade (GDS3, GB2), Dawson (MMW9), Floyd
(MD3), Gilmer (GB2), Jasper (GB2), Rabun (GBi), Union (FSCA), Walker (GDS3),
Whitfield (GB2).
Uncommon in the mountains and
upper Piedmont, mostly at shallow ponds. [May 14 - Aug 29]
Enallagma basidens
Calvert, 1902.
Double-striped Bluet. State: W&M (1996*), Donnelly (2004c).
Bartow (UG), Ben Hill (WMC), Carroll (GB2), Chattooga (MD4), Clarke
(FSCA, UG), Dade (GDS3), Decatur (FSCA), Floyd (MD4), Franklin (WMC), Gwinnett
(PA), Hall (FSCA, MW9), Haralson (GB2), Irwin (GB3), Jones (WMC), Lanier (MD4),
Long (GB3), Monroe (GB3), Morgan (MV3), Murray (UG), Paulding (GB2), Polk
(GB2), Rabun (FSCA), Screven (FSCA), Taliaferro (FSCA), Walker (GB2, MD3),
Wilcox (GB2).
Common above the fall line,
uncommon below, and rare in the extreme southeast at slow streams, ponds, and
lakes. [Apr 19 - Oct 23]
Enallagma civile
(Hagen, 1861). Familiar Bluet. State: W&M (1996), Donnelly (2004c).
Baker (GB2), Berrien (MD4), Catoosa (UN), Cherokee
(GB3), Clarke (UG), Clay (GB3), Clinch (GB3), Coffee (RB), Crisp (GB2, GB4),
Dade (GDS3, GB2), Dawson (MW9), Floyd (MD3), Gilmer (GB3), Glynn (GB2, GB4),
Laurens (GB3), Lee (RB), McDuffie (Wi34*), Meriwether (GB2), Morgan (MV3),
Murray (KT), Paulding (GB2), Rabun
(GB4), Richmond (GB1), Union (FSCA), Walker (GDS3), White (GB3), Whitfield
(GB2).
Common
throughout the state at any water habitat, especially ponds. [Apr 3 -
Nov 29]
Enallagma coecum
(Hagen)*. Purple Bluet. State: Donnelly (2004c).
Atkinson (GB4, MD4), Clinch (TD*), Early
(GB4).
Rare in extreme
south Georgia where it reaches its northern range limit. Prefers tannic slow
streams. The Clinch County record is: Suwanee R. at Fargo, 30°41.0’N
82°33.6’WMC, 2 Apr 2002, 1 pr tandem. Coll. T. Donnelly. The Early County record is on a small
unnamed stream with numerous males found in both 2004 and early 2005. Dunkle
(1992) reports it across northern Florida, so it should be found in additional
counties in southern Georgia. [Apr 2 - Oct 23]
Enallagma concisum
Williamson, 1922. Cherry Bluet. State: W&M (1996), Donnelly (2004c).
Ben Hill (WMC), Bulloch (DP), Charlton (UG), Fayette
(UG), McDuffie (UN, Wi34*), Telfair (GB4, MD4), Treutlen (MD4), Ware (MV3, UG),
Wheeler (DP).
Uncommon to rare in
southeastern Georgia; rare just above the fall line, in shallow ponds. [Apr 4 - Sep 27]
Enallagma daeckii
(Calvert, 1903). Attenuated Bluet. State: W&M (1996), Donnelly (2004c).
Charlton (UG), Coffee (DP), Colquitt (UG), Crawford
(UN), Early (MV3), Lee (Ro24*), Lowndes (WMC), Morgan (DP), Murray (MD4),
Taylor (MD4), Telfair (GB4, DP), Towns (MD4), Wayne (DP), Wheeler (DP).
Uncommon below the fall line
at ponds and lakes; rare above the fall line. [Apr 11 - Sep 10]
Enallagma davisi Westfall,
1943. Sandhill Bluet.
State: W&M (1996*), Donnelly (2004c).
Taylor (FSCA, GB3), Telfair (GB 2005).
Should be rare at sandy lakes
and their outflows in the fall line sandhills or elsewhere in the coastal
plain. It is known from only two
counties so far. Dunkle (1992) reports it as “uncommon at sand bottomed lakes
across north Florida”. [Mar 14 - Apr 27]
Enallagma divagans
Selys, 1876. Turquoise Bluet. State: W&M (1996*), Donnelly (2004c).
Banks (GB1), Charlton (FSCA), Chatham (UG),
Chattooga (GB2), Clarke (UG), Clayton (UG), Crawford (UG), Dade (GDS3), Dawson
(MW9), Fayette (UG), Floyd (MD4), Franklin (WMC), Gilmer (WMC), Greene (UG),
Haralson (WMC), Jones (MV3, WMC), Monroe (MV3), Morgan (MV3, UN), Peach (UG),
Rabun (FSCA), Richmond (UN), Talbot (JD), Taylor (RB3, UG), Walker (GDS3),
Walton (UG).
Uncommon at streams above the
fall line, and very rare below. [Apr 18 - Jul 6].
Enallagma doubledayi
(Selys, 1850). Atlantic Bluet. State: W&M (1996), Donnelly (2004c).
Baker (FSCA), Berrien (FS2), Clarke (UG), Clinch
(GB3), Dooly (By31), Early (GB4), Effingham (UN), Gwinnett (PA), Laurens (GB2),
Lee (Ro24*), Long (GB3, WMC). McDuffie (Wi34), Montgomery (MD4), Morgan (DP),
Tattnall (GB4), Wheeler (DP).
Fairly common south of the
upper Piedmont at ponds, lakes and slow rivers. It has probably been
overlooked. [Apr 8 - Oct 29]
Enallagma dubium
Root, 1924. Burgundy Bluet. State: N&H (1929), Byers (1930), W&M (1996), Donnelly (2004c).
Bacon (MW41), Ben Hill (WMC), Charlton (UG), Crawford (UG), Dade (GDS3), Dooly (UG), Lee
(Ro24*), Lowndes (MD4), McDuffie (UN, Wi34), Richmond (UN), Tattnall (GB4),
Taylor (GB3), Treutlen (GB3), Walker (GDS3), Wayne (DP).
Uncommon at lakes and ponds
below the fall line, rare above. [Apr 4 - Sep 28]
Enallagma durum
(Hagen, 1861). Big Bluet. State: W&M(1996*), Donnelly (2004c).
Baker (FSCA).
Should be in more
counties in extreme southeast near coast, but known only from one county in the
southwest. Dunkle (1992) reports it from south of Tallahassee, Florida to north
of Jacksonville in brackish habitat. [No date information].
Enallagma exsulans
(Hagen, 1861). Stream Bluet. State: W&M (1996*), Donnelly (2004c).
Chattooga (GDS3), Dade (GDS3, GB2), Floyd (MD3),
Gordon (UN), Haralson (GB2), Harris (GB1), Polk (GB2), Talbot / Upson (GB2),
Walker (GDS3).
A Clinch County record previously reported (Donnelly
2004c) was actually a juvenile E. weewa
(Donnelly, pers. comm.). Fairly common in streams and rivers to just below
the fall line. [Apr 29 - Sep 11]
Enallagma geminatum
Kellicott, 1895. Skimming Bluet. State: W&M (1996), Donnelly (2004c).
Bartow (UG, UN), Ben Hill (GB2), Bulloch (DP), Burke
(GB3, Wi34), Carroll (GB2), Coffee (DP), Crawford (UN), Dade (GDS3), Dawson
(MW9), Early (GB3), Emanuel (GB4, KT), Floyd (MD4), Franklin (WMC), Gordon
(MD4), Haralson (GB2), Houston (UG, UN), Jones (WMC), Laurens (WMC), Lee
(Ro24*), McDuffie (Wi34), Paulding (GB3), Polk (GB1), Richmond (UN), Tattnall
(WM95a, JD), Taylor (GB3), Walker (GB2), Wheeler (DP), White (GB4).
Fairly common throughout the
state at lakes and ponds; rare below the fall line in slow streams. [Mar 17 - Oct 3].
Enallagma hageni
(Walsh, 1863). Hagen's Bluet. State: W&M (1996*), Donnelly (2004c).
Rabun (FSCA).
Known only from the extreme
northeast part of Georgia: Rabun County, pond 1.1 mi. E of Satolah. In the FSCA there are 23 males and 1 female
from the same locality collected in 1972. This common northern species probably
reaches its southern limit in north Georgia. [Jun 12 - Aug 10].
Enallagma pallidum Root,
1923. Pale Bluet.
State: W&M (1996*), Donnelly (2004c).
Appling (FSCA), Coffee (DP), Echols (WMC), Effingham
(GB2), Emanuel (GB1), Tattnall (GB4).
Rare in the eastern coastal
plain at lakes and ponds; very rarely found in slow streams. [May 11 - Jun 13].
Enallagma pollutum
(Hagen, 1861). Florida Bluet. State: W&M (1996*), Donnelly (2004c).
Charlton (DP).
It is known only from one
record in southeast Georgia, but since Byers (1927c, 1930) reports it from Leon
County Florida, it should be found in southwest Georgia also. [No date information].
Enallagma signatum
(Hagen, 1861). Orange Bluet. State: Hagen (1861*, 1863, 1874), Banks (1892), Calvert (1893), Muttkowski (1910), W&M (1996), Donnelly (2004c).
Atkinson (DP), Bartow (MD4), Ben Hill (GB2, WMC),
Brantley (GB4), Brooks (GB1), Burke (Wi34), Butts (WMC), Carroll (GB2, WMC), Catoosa
(GB2), Charlton (DP), Clay (GB3), Cook (GB1, GB2), Dade (GDS3, GB2), Decatur (By27c), Early (GB3, MV3), Emanuel
(GB1), Fannin (GB4), Fayette (KT), Floyd (MD4), Franklin (WMC), Fulton (HS4),
Gilmer (GB3), Hancock (MD4), Harris (GB1, GB3, MV3), Irwin (GB3), Jeff Davis
(JF3), Jones (MV3), Lanier (MD4), Laurens (By31), Lee (Ro24), Long (GB3, DP),
Lowndes (MD4), Lumpkin (GB1, MV3), McDuffie (Wi34), Meriwether (GB2), Monroe
(GBi, GB2, GB3), Montgomery (GB2), Morgan (UG), Murray (SR), Newton (UG),
Paulding (GB), Polk (GB), Rabun (GBi), Richmond (UN), Sumter (RB), Talbot /
Upson (GB2), Taliaferro (GB2), Tattnall (GB1), Taylor (GB3), Tift (MD3), Toombs
(GB1), Treutlen (GB1), Union (GB3), Walker (GDS3, GB2), Ware (MV3), Washington
(MD4), Wayne (GB3,DP).
Common throughout the state at
ponds, lakes and streams; uncommon at larger rivers. [Apr 3 - Nov 4].
Enallagma traviatum
traviatum Selys, 1876. Slender Bluet. State: W&M (1996), Donnelly (2004c).
Ben Hill (WMC), Clarke (UG), Cobb (GB3), Crawford
(UG), Dade (GDS3), Fannin (GB4), Floyd (MD4), Franklin (WMC), Gilmer (GB2,
GB3), Jones (WMC), Monroe (GB3, MD3), Montgomery (GB2), Rabun (TD73*), Union
(GB3), Walker (GDS3), Wilkinson (DP).
Uncommon throughout the state
at lakes and ponds, but rare or absent from the deep south and near the coast. [May 15 - Aug 15].
Enallagma vesperum
Calvert, 1919. Vesper Bluet. State: W&M (1996), Donnelly (2004c).
Brantley (DP), Brooks (GB1), Bulloch (DP), Early
(MV3), Floyd (MD4), Harris (GB1), Long (GB3), Lowndes (GB4, MD4), McDuffie
(Wi34*), Richmond (UG, UN), Tattnall (KT, SK, JD, WM95a), Taylor (GB3).
Uncommon throughout state at
ponds and lakes with lily pads, but probably overlooked due to its crepuscular
habits. [Apr 1
- Nov 3].
Enallagma weewa
Byers, 1927. Blackwater Bluet. State: Byers (1927b*, 1930), N&H (1929), W&M (1996), Donnelly
(2004c).
Atkinson (GB4), Berrien (GB3), Brantley (GB4, DP),
Charlton (DP, UG), Clinch (TD), Coffee (GB3), Crawford (UG), Evans (DP), Greene
(MD4), Lanier (UG, UN), Macon (By27c), McDuffie (UN, Wi34), Oglethorpe (By27c),
Richmond (GBi, UN), Tift (GB3), Wayne (DP).
Uncommon below the fall line
in tannic rivers and streams. [May 15 - Nov 4]
Ischnura hastata
(Say, 1839). (Anomalagrion hastatum of some authors). Citrine Forktail. State: Hagen (1874),
W&M (1996), Donnelly (2004c).
Atkinson (DP), Bartow (UG), Brantley (BB), Brooks
(FSCA, GB1), Bulloch (DP), Burke (Wi34), Calhoun (MV3), Camden (UN), Carroll
(GB, GB2), Charlton (Br14, PA, WMC), Chatham (Ha61*), Chattooga (GDS3),
Cherokee (GB), Clarke (UG), Clinch (GB3, WMC), Cobb (GB), Coffee (GB3, GB4),
Columbia (UN), Cook (MD3), Crawford (UG), Dade (GDS3, GB2), Dawson (WMC),
Decatur (UG), Early (MV3), Echols (GB4), Emanuel (GB1), Fannin (GB3), Floyd
(MD4), Franklin (UG, WMC), Gilmer (GB3), Glynn (WMC), Gwinnett (By31, PA),
Habersham (UG), Haralson (WMC), Harris (MV3), Irwin (RB), Jones (WMC), Laurens
(GB1, WMC), Lee (Ro24, RB3, UG), Long (WMC), Lowndes (UG, WMC), Lumpkin (GB1),
McDuffie (UN, Wi34), McIntosh (MV3, DP, UG), Meriwether (GB2), Montgomery (GB1),
Morgan (DP), Murray (GB3, SK), Polk (GB), Rabun (GB3), Richmond (UN), Stephens
(GB2), Tattnall (MV3, WMC), Taylor (GB3, UG, UN), Telfair (WMC), Toombs (KT),
Towns (KT), Walker (GDS3, GB2), Wayne (OO, DP), Wheeler (GB1, WMC), Wilcox
(GB2), Worth (WMC).
Common throughout the state in
grassy areas along ponds, lakes, streams, and rivers. [Mar 13 - Jan 6].
Ischnura kellicotti
Williamson, 1898. Lilypad Forktail. State: W&M (1996), Donnelly (2004c).
Atkinson (RB), Berrien (GB1), Brooks (GB1), Bryan
(GB1), Burke (Wi34*), Butts (MD4), Charlton (OO), Chatham (GB2), Colquitt
(MD4), Cook (GB3), Dodge (GB4), Early (GB2), Echols (WMC), Jeff Davis (MD4),
Long (DP), Lowndes (GB4, MD3), Meriwether (MD4), Montgomery (GB1), Screven
(DP), Sumter (GB2), Tattnall (KT, WM95a), Tift (RB), Treutlen (MD4), Turner
(GB3), Wayne (OO), Wheeler (WMC), Wilkinson (GB3).
Fairly common at ponds and
lakes with lily pads below the fall line. [Mar 21 - Nov 3].
Ischnura posita
(Hagen, 1861). Fragile Forktail. State: Hagen (1874), Banks (1892) as Nehalennia posita, Calvert
(1893), Muttkowski (1910), W&M (1996),
Donnelly (2004c).
Appling (GB), Bartow (GB1, UG), Brantley (BB),
Brooks (GB1), Bulloch (DP), Burke (Wi34), Butts (MD4), Calhoun (MV3), Candler
(GB1), Carroll (GB2, WMC), Catoosa (GB), Chatham (Ha61*, Ha63, Se76), Chattooga
(GB, SR), Clarke (UG), Clayton (GB), Cobb (GB), Coffee (DP), Cook (GB1, GB2),
Crawford (UG), Dade (GDS3, GB2), Dawson (WMC), Dooly (By31), Douglas (GB),
Early (MV3), Echols (GB4), Effingham (GB2), Emanuel (GB1), Fayette (KT, UG),
Floyd (GB1), Franklin (UG, WMC), Fulton (HS4), Gilmer (WMC), Grady (GB1),
Gwinnett (GB, GB2, PA), Habersham (GBi, UN), Haralson (WMC), Harris (MV3),
Irwin (GB1), Jasper (GB2), Jeff Davis (MD4), Jones (MV3, WMC), Lanier (MD4),
Laurens (GB1), Lee (Ro24, UG), Liberty (UN), Long (DP), Lowndes UG), Lumpkin
(GB1), McDuffie (Wi34), McIntosh (MV3), Meriwether (GB1), Monroe (GB2),
Montgomery (GB1), Morgan (MV3, DP, UG), Murray (SK, SR, WMC), Newton (UG), Oglethorpe (UG), Paulding (GB), Polk (GB), Pulaski
(MD4), Putnam (UG), Rabun (GBi), Richmond (UG, UN), Rockdale (GB), Stephens
(GB2), Sumter (RB), Tattnall (GB4, MV3, WMC), Taylor (UG, UN), Telfair (WMC),
Thomas (GB1), Tift (RB), Toombs (GB1, WMC), Truetlen (GB1), Turner (GB3),
Walker (GDS3, GB2), Ware (UG), Wayne (DP), Wheeler (GB1, WMC), White (GB3,
GB4), Whitfield (Ha61*, Ha63, Se76), Wilcox (GB2), Wilkinson (DP), Worth (WMC).
Common throughout the state
near ponds and lakes, and uncommon along streams and rivers. [Mar 1 - Jan 6].
Ischnura prognata
(Hagen, 1861). Furtive Forktail. State: W&M (1996), Donnelly (2004c).
Glynn (GB3), Lee (Ro24*), Liberty (DS), Long (DS),
Tattnall (WM95a), Telfair (WMC), Toombs (WMC), Wayne (DP).
Rare in the southern and
eastern coastal plain in seeps and swampy areas. [Mar 14 - Sep 7].
Ischnura ramburii
(Selys, 1850). Rambur's Forktail. State: Hagen (1874), Cuyler (1989b), W&M (1996), Donnelly (2004c).
Appling (GB4), Baker (MV3), Ben Hill (GB2, GB4),
Berrien (FS2), Bibb (UG), Brooks (GB1), Bryan (GB, GB1, UN), Carroll (WMC),
Charlton (OO, DP), Chatham (GB), Clinch (TD, WMC), Coffee (GB3, RB), Colquitt
(MD3), Crisp (GB2, GB4), DeKalb (UG), Effingham (GB2, UN), Evans (GB1), Floyd
(MD4), Glynn (GB1, GB4), Grady (GB1), Greene (GB), Gwinnett (PA), Haralson
(WMC, GB2), Irwin (GB3, RB), Jasper (GB2), Jefferson (GB2), Jeff Davis (MD4),
Lanier (GB3, MD4), Laurens (GB1), Lee (Ro24, RB), Liberty (UG), Long (GB3, BB),
Lowndes (MD4), McDuffie (Wi34), McIntosh (MV3, UG), Montgomery (GB1, GB2),
Muscogee (GB), Polk (GB), Seminole (GB), Stewart (GB), Taliaferro (GB2),
Tattnall (GB4, WMC), Telfair (WMC), Tift (RB), Toombs (GB1), Treutlen (MD4),
Turner (WMC), Union (GB1), Walker
(GDS3, MD4), Ware (MV3), Wheeler (DP), Whitfield (Ha61*, Ha63), Worth (WMC).
Common throughout the state at
lakes, rivers, and streams; often abundant at ponds. [Mar 21 - Dec 22].
Ischnura verticalis (Say,
1839). Eastern Forktail.
State: Selys (1876*), Calvert (1893, 1903a), Muttkowski
(1910), N&H (1929), W&M (1996), Donnelly (2004c).
Burke (By31), Chattooga (GDS3), Clarke (UG), Clayton
(UG), Dade (GB2), Dawson (MW9), DeKalb (UG), Franklin (WMC), Newton (UG), Rabun
(GBi, GB4) Stephens (UN), Union (GB3), Walker (GDS3).
Uncommon in north Georgia at
lakes; fairly common at shallow ponds; and rare at the same habitats in the
Piedmont. [May
23 - Sep 24].
Nehalennia gracilis
Morse, 1895. Sphagnum Sprite. State: W&M (1996), Donnelly (2004c).
Long (GB3), White (SD83*).
Only known from two records
but at opposite ends of the state; possibly rare throughout in small ponds. Donnelly (2004c) shows it
from scattered localities across northern Florida, central Alabama, and western
North Carolina, so we expect it to be found in more localities in Georgia. [May
8 - May 25].
Nehalennia
integricollis Calvert, 1913. Southern Sprite. State: W&M (1996), Donnelly (2004c).
Charlton (DP, UN), Clinch (WMC), Coweta (FSCA),
Early (FSCA), Fannin (GB3), Laurens (GB4), Lee (Ro24), McDuffie (Wi34), Morgan
(DP), Rabun (FSCA), Richmond (SD), Screven (DP), Telfair (MD4), Thomas (Ca13*; type locality: Thomasville), Walker (MD3), Ware (RB,
UG), Wheeler (DP).
Uncommon throughout the state
in marshy or grassy pond and lake edges. [Apr 5 - Sep 28].
Telebasis byersi
Westfall, 1957. Duckweed Firetail. State: W&M (1996), Donnelly (2004c).
Appling (GB), Bryan (DS), McIntosh (GBi90*).
Rare in eastern coastal plain
in slow margins of rivers, probably overlooked. [Jul 9 - Jul 31].
ANISOPTERA
Tachopteryx thoreyi
(Hagen in Selys, 1858). Gray Petaltail. State: Louton (1982), NW&M
(2000), Donnelly (2004b).
Baldwin (SP), Bartow (GB1, WMC), Butts
(WMC), Carroll (GB2), Cherokee (UG), Clarke (UG), Dawson (GB4), Floyd (MD4),
Forsyth (RT), Franklin (WMC), Jackson (UG), Jefferson (UG), Long (GB3), Lumpkin
(UG), McIntosh (UG), Monroe (GB4), Morgan (FSCA), Murray (WMC), Paulding
(GB), Rabun (Da11*, UG), Talbot (JD),
Walker (MD4), Wayne (DS, GB3), White (UG).
Fairly common at
seeps and associated rivers and streams above the fall line and scattered
below, probably due to a paucity of suitable habitat. [Apr 12 - Jul
18].
Aeshna umbrosa umbrosa
Walker, 1908. Shadow Darner. State: NW&M (2000), Donnelly (2004b).
Cobb (GB, GB2), Fannin (SD83*, UG), Floyd (GB2),
Fulton (UG), Glascock (JF), Hall (D8), Jackson (UG), Murray (GB3), Oconee (UG),
Pickens (UN), Towns (D8, UG).
Uncommon from the middle
Piedmont north in or near streams and small rivers, rare at ponds. [Aug 21 - Dec 7].
Anax junius
(Drury, 1770). Common Green Darner.
State: Hagen (1861*, 1863, 1874, 1875, 1890a), N&W (1955), NW&M
(2000), Donnelly (2004b).
Bacon (GB1), Baker (GB2), Baldwin (SP),
Bartow (GB1), Berrien (MD4), Bibb (UG), Brantley (BB), Calhoun (MV3), Camden
(GB), Carroll (GB), Charlton (OO, PA), Chattooga (GB2, SR), Cherokee (GB),
Clarke (UG), Clayton (GB), Clinch (UG), Cobb (GB), Coffee (GB3), Colquitt
(MD4), Coweta (GB), Dade (GDS3), Decatur (UG), DeKalb (GB, UG), Early (MV3),
Emanuel (UG), Evans (WMC), Fannin (JB3), Floyd (MD3), Forsyth (GB), Glynn
(WMC), Gwinnett (PA), Harris (MV3), Johnson (SD), Jones (MV3), Lamar (GB),
Laurens (GB, PA), Lee (Ro24, RB3), Lincoln (UG), Long (WMC), Lumpkin (MV3),
McDuffie (Wi34), McIntosh (GB, MV3), Meriwether (GB2, MV3), Mitchell (UG),
Monroe (MV3), Morgan (MV3), Murray (SK), Newton (MV3), Oglethorpe (UG),
Paulding (GB), Peach (GB, UN), Rabun (GBi, UN), Richmond (SD, UN), Rockdale (FS2), Seminole (GB), Stewart (GB),
Talbot / Upson (GB2), Tattnall (MV3, WMC), Taylor (S), Tift (LT), Toombs (SD),
Union (SD), Walker (GB2), Ware (MV3, RB), Wayne (OO), White (S, WMC), Wilcox
(GB2), Wilkes (UG).
Common statewide
at ponds, lakes, and marshes, but can be found almost anywhere, including along
the coast as a migrant. It probably flies year round in warm winters. [Jan 28 - Jan
6].
Anax longipes
Hagen, 1861. Comet Darner. State: Hagen (1861*, 1863,
1866, 1874, 1875, 1890a), Banks (1892), Calvert (1906), Muttkowski (1910) as concolor, Needham & Westfall (1955), Ries & Cruden (1966), NW&M (2000),
Donnelly (2004b).
Appling (GB), Baldwin (SP), Bartow
(GB1), Bibb (UG), Bryan (DS), Bulloch (AH1, AH3), Burke (DS), Chattooga (GB2),
Cobb (GB3), DeKalb (GB), Floyd (MD4), Laurens (GB3), Lee (Ro24)
Liberty (UG), Long (DS), Rockdale (FS2),
Talbot (GB1), Tattnall (DS), Walker (GDS3), Wheeler (DP).
Uncommon across the state at
small ponds and sterile shallow pits. [Mar 27 - Sep 4].
Basiaeschna janata
(Say, 1839). Springtime Darner. State: Hagen (1863*, 1874), Montgomery (1947), NW&M (2000),
Donnelly (2004b).
Bartow (GB2, UG, UN), Burke (UG), Chattooga (SR),
Fannin (MV3), Floyd (MD4), Gordon (SR), Harris (MV3), Jones (MV3), Meriwether
(MV3), Monroe (MV3), Morgan (MV3), Murray (Lo82, SR, UN, WMC), Paulding (GB3),
Richmond (FSCA), Taylor (GB4), Toombs (WMC), White (GB4).
Uncommon to fairly common at
streams above the fall line, rare below the fall line or above the fall line
along large rivers and ponds near streams. [Mar 20 - May 16].
Boyeria grafiana
Williamson, 1907. Ocellated Darner. State: NW&M (2000), Donnelly (2004b).
Lumpkin (SD83*), Murray (GB3, GB4), White (D8, JD).
Rare at fast cold streams in
the northeast corner of the state. [Jul 31 - Sep 23].
Boyeria vinosa
(Say, 1839). Fawn Darner.
State: Hagen (1863*, 1874) as Aeschna
quadriguttata, (1875), Banks (1892) as Neuraeschna vinosa,
N&W (1955), NW&M (2000), Donnelly (2004b).
Baldwin (SP), Bibb (UG), Brantley (DP), Burke (By31,
Wi34), Carroll (GB), Chattooga (GB4, UG), Cherokee (GB2, Lo82, SK, UN), Clarke
(UG), Cobb (GB1, GB2, GWQ71), Columbia (Lo82), Coweta (GB, GWQ71), Emanuel
(GB4), Fannin (JB3), Floyd (GB2), Forsyth (GWQ71), Fulton (GWQ71, UG), Gilmer
(Lo82, UN), Gordon (GB2), Gwinnett (GWQ71), Habersham (GWQ71), Haralson (GB2), Heard (GB2), Houston (SD), Jefferson
(SD, Wi34), Lumpkin (GWQ71), McDuffie (Wi34), Meriwether (SD), Murray (GB3,
Lo82), Muscogee / Chattahoochee (GWQ71), Oglethorpe (MD4), Paulding (MD4), Polk
(GB2), Quitman (GWQ71), Rabun (SD), Richmond (Lo82), Rockdale (FS2), Toombs /
Tattnall (Lo82), Talbot / Upson (GB2), Troup (GWQ71), Union (SD, JD, Wi34),
White (GWQ71, Wi34).
Mostly uncommon
at streams and rivers, but fairly common above the fall line. [May 22 - Oct 4].
Coryphaeschna ingens
(Rambur, 1842). Regal Darner. State: Hagen (1874*) as Aeschna abboti,
(1875), Kirby (1890). Banks (1892) as Aeshna
ingens, Calvert (1903b, 1906), Williamson (1903a), Martin (1908), Muttkowski (1910), N&H (1929), Byers (1930),
N&W (1955), NW&M (2000), Donnelly (2004b).
Baker (By31), Calhoun (MV3), Charlton (SD, UN),
Clinch (DP, UG), Early (GB2), Glynn (By31), Lee (Ro24) McIntosh (MV3, UG),
Monroe (MV3), Wheeler (DP).
Uncommon to rare below the
fall line; very rare above, at ponds and lakes. [Apr 6 - Sep 23].
Epiaeschna heros
(Fabricius, 1798). Swamp Darner. State: Hagen (1874*, 1875), N&W (1955), NW&M (2000), Donnelly
(2004b).
Appling (GB2), Baker (MV3), Baldwin (SP), Bartow
(GB2), Bibb (UG), Brantley (BB), Burke (By31), Calhoun (MV3), Charlton (Br14,
OO, UG), Cherokee (GB3), Clarke (UG), Clinch (UG), Cobb (GB1), Coffee (GB3),
Dade (GDS3), DeKalb (UG), Dooly (GB2), Dougherty (UG), Early (GB3), Evans (GB),
Fulton (AH2, UG, UN), Glynn (By31, GB4), Gwinnett (PA), Heard (GB1), Jones
(MV3), Laurens (GB2), Liberty (UG), Long (GB3, WMC), Lowndes (WMC), Lumpkin
(MV3), McIntosh (GB, MV3), Meriwether (MV3), Monroe (MV3), Morgan (MV3), Murray
(GB2, KT, WMC), Pike (UN), Putnam
(MV3), Rockdale (FS2), Tattnall (MV3, WMC), Toombs (WMC), Walker (GDS3),
Ware (MV3), Wayne (OO), White (SD), Wilkinson (DP).
Common throughout the state in
forested habitat along rivers and streams; uncommon at forested or swampy
ponds. [Mar 21
- Nov 2].
Gomphaeschna antilope
(Hagen, 1874).Taper-tailed Darner. State: Hagen (1874*), N&W (1955), NW&M (2000), Donnelly
(2004b).
Bibb (UG), Camden (DG40), Charlton (DG40,OO), Clarke
(UG), Coffee (GB3), Coweta (FSCA), DeKalb (DG40, UN), Elbert (GB3), Evans (KT),
Fulton (UG), Jones (FSCA), Liberty (UG) Lincoln (GB3), McIntosh (UG), Taylor
(SD).
Rare throughout the state near
streams, rivers, or swamps but absent from extreme north. [Mar 21 - May 15].
Gomphaeschna
furcillata (Say, 1839). Harlequin Darner. State: Hagen (1874*, 1875), Banks (1892), Muttkowski (1910), N&H (1929), N&W (1955), NW&M (2000), Donnelly (2004b).
Baldwin (SP), Bibb (UG), Brantley (BB), Burke (GB3),
Cobb (GB1), Coffee (GB3), Crawford / Bibb (SD), Early (GB4), Effingham (GB4),
Harris (MV3), Pierce (UN), Thomas (DG40).
Rare throughout the state at
or near swamps. [Mar
17 - Apr 15].
Gynacantha nervosa Rambur,
1842. Twilight Darner.
State: NW&M (2000), Donnelly (2004b).
Charlton (GB4, UG), Glynn (UG), Thomas (Ko60*).
Rare in the extreme southeast
near small ponds along forested rivers. [Only date recorded: Nov 4].
Nasiaeschna
pentacantha (Rambur, 1842). Cyrano Darner. State: N&W (1955), NW&M (2000), Donnelly
(2004b).
Baker (UG), Charlton (UN), Coffee (DP), Glynn (By31*), Habersham (GBi), Hancock (SD), Jasper (GB2), Laurens (GB1), Lowndes (WMC), Toombs (KT), Wayne (By31).
Scattered records across the state; probably uncommon, along slow rivers
and streams. [Apr 1 - Jul 31]
Triacanthagyna trifida
(Rambur, 1842). Phantom Darner. State: N&W (1955), NW&M (2000), Donnelly (2004b).
Camden (GB4), Charlton (SD, GB4, Wi23a*), Glynn (UG,
Wi23a*), Liberty (DS2, GB3), Ware (WMC).
Fairly common in the southeast
and along the coast; easily overlooked due to crepuscular nature; near small
ponds along forested rivers. [Oct 11 - Dec 14].
Aphylla williamsoni (Gloyd,
1936). Two-striped Forceptail. State: NW&M (2000), Donnelly (2004b).
Berrien (GB), Bryan (GB1), Bulloch
(AH1), Grady (GB1), Crisp (GB2), Dodge (GB1), Harris (GB), Jasper (GB2), Lamar
(GB), Laurens (GB), Liberty (DS), Long (DS), Miller (GB), Rockdale (FS2),
Seminole (GB), Talbot / Upson (GB2), Wilcox (GB2).
Recently discovered in Georgia, this
species is common at ponds and lakes below
the fall line; rare but slowly expanding north of the fall line into the middle Piedmont. [Jul 3 - Oct 4].
Arigomphus pallidus
(Rambur, 1842). Gray-green Clubtail. State: Selys (1858*), Hagen (1861, 1863, 1874, 1875), also as G.
pilipes, Banks (1892), Muttkowski (1910), N&H (1929), Byers (1930),
N&W (1955), NW&M (2000), Donnelly (2004b).
Baker (UG), Bryan (DS), Clinch (WMC), Coffee (MD4),
Early (FSCA, GB2), Lanier (GB3), Long (DS), Lowndes (MD4, WMC), Tattnall (DS),
Thomas (Wi14).
Fairly common in the southern
and eastern coastal plain at ponds, lakes, and the slow edges of streams and
rivers. [Apr 27
- Jul 24].
Arigomphus villosipes
(Selys, 1854). Unicorn Clubtail. State: Donnelly (2004b).
Dade (WM04*), Walker (MD4 photo).
Very rare in the extreme
northwestern section of the state; two records at ponds. The Dade County record
is: Trenton,
Cloudland Canyon State Park, pond, N 34° 49.58’ W 85° 28.61’, 23 May 2003, 1
male, Coll. Mike Thomas [May 23 - Jul 4].
Dromogomphus armatus
Selys, 1854. Southeastern Spinyleg. State: Selys (1878a*, Banks (1892), Muttkowski (1910), N&H (1929),
N&W (1955), Louton (1982), Westfall & Tennessen (1979), NW&M
(2000), Donnelly (2004b).
Burke (By31, Wi34), Early (SK4), Jefferson (FSCA).
Rare below the fall line at
forested mucky seeps or nearby. Dunkle (1992)
reported scattered records across north Florida, so it should be found at additional Georgia
locations. [Aug 19 - Sep 17].
Dromogomphus spinosus Selys, 1854. Black-shouldered
Spinyleg. State: Selys
(1858*), Hagen (1861, 1863, 1874, 1875), Banks (1892), Calvert (1893), N&W
(1955), Westfall & Tennessen (1979), NW&M (2000), Donnelly (2004b).
Baker (JF4), Baldwin (SD, SP), Burke
(SD), Calhoun (UG), Catoosa (GB, Lo82, UN), Cherokee (GB2), Cobb (GB2),
Crawford (GB), Crisp (GB2), Dade (Lo82, UN), Decatur (UG), Douglas (GB), Early
(GB3, GB4), Emanuel (SD, GB1), Floyd (By31, GB1), Gilmer (GB3), Gwinnett (GB,
JF), Hancock / Warren (SD), Haralson (GB2), Jefferson (GB1), Laurens (By31),
Lee (GBi), McDuffie / Warren (SD), Monroe (GB3), Murray (Lo82), Paulding (GB2),
Polk (GB), Randolph (JF4), Rockdale (FS2), Stephens
(GB2), Talbot / Upson (GB2), Tattnall (GB1), Telfair (MD3), Truetlen (GB1), Whitfield (MD3), Wilkinson (By31).
Common throughout
the state along streams and rivers except rare at lakes or along the coast. [May 19 - Sep
20].
Dromogomphus spoliatus (Hagen in Selys, 1858). Flag-tailed Spinyleg. State: NW&M
(2000), Donnelly (2004b).
Chattooga (GB2), Catoosa (GB), Floyd
(GB, MD3).
Rare at rivers in
the extreme northwestern section of the state; very rarely at ponds. It was
originally reported by Montgomery (1947), which was based on a misidentified
specimen (Tennessen, 1979). [Jun 20 -
Sep 21].
Erpetogomphus designatus Hagen in Selys, 1858. Eastern Ringtail. State: N&W
(1955*), Louton (1982), NW&M (2000), Donnelly (2004b).
Baldwin (SP), Chattooga (UG), Clarke
(UG), Decatur (UG), Houston (SD, RG94b, JD, UN), Laurens (GB3), Long (GB3),
Talbot / Upson (GB2), Tattnall (DS, GB3), Toombs (GB1), Twiggs (JD).
Uncommon on large
rivers throughout the state, except absent from the immediate coastline. [Jun 13 - Oct
27].
Gomphus (Gomphurus) consanguis (Selys, 1879). Cherokee Clubtail. State: NW&M
(2000), Donnelly (2004b).
Chattooga
(GBi03, GDS3, GB2), Floyd (GB4), Walker (GBi83b*, GBi03, SD83*, GDS3).
Rare to locally uncommon on small streams, often spring-fed, in the
northwestern section of Georgia. [May 23 - Jun 20].
Gomphus (Gomphurus) dilatatus Rambur, 1842. Blackwater Clubtail. State: Hagen
(1861*, 1863, 1874, 1875), Banks (1892), Muttkowski (1910), Calvert (1921,
1922), N&H (1929), Byers (1930), N&W (1955), Westfall (1974), NW&M
(2000), Donnelly (2004b).
Baldwin (SP),
Bryan (DS), Decatur (FSCA), Early (GB3), Echols (UN), Hall (UN), Houston (SD),
Monroe (GB3, MV3), Murray (Lo82), Newton (MV3), Schley (UG), Tattnall (DS),
Telfair (MD3), Upson (UG).
Uncommon along rivers and streams below the fall line, rare above. [Apr 11 - Jul
13].
Gomphus (Gomphurus) hybridus Williamson,1902. Cocoa Clubtail. State: NW&M
(2000), Donnelly (2004b).
Baldwin (SP),
Crawford / Taylor (SD), Gilmer (Lo82), Jeff Davis (Lo82*, UN), Laurens (GB4),
Long (GB3), Monroe (SD), Montgomery (Lo82*, UN), Murray (SD), Tattnall (KT, WM95a),
Taylor / Upson (SD), Toombs (KT, SK, WMC), Twiggs (GB4), Wayne (GB3).
Rare along larger rivers and streams across the state. It should be found
in the southwestern part of the state, since Westfall (1953 ) reports it from
nearby Liberty and Gadsden counties in Florida. [Apr 1 - May 15].
Gomphus (Gomphurus) lineatifrons Calvert, 1921. Splendid Clubtail. State: NW&M
(2000), Donnelly (2004b).
Chattooga (GDS3), Cherokee (Lo82*), Dade
(GDS3, Lo82*, UN). Floyd (MD3), Gilmer (Lo82*), Walker (GDS3), Whitfield (MD4).
Uncommon
to rare along rivers and large streams in the extreme northwestern corner of
the state. [May
23 - Jun 26].
Gomphus
(Gomphurus) rogersi (Gloyd, 1936). Sable Clubtail. State: NW&M (2000),
Donnelly (2004b).
Fannin (SD, GB3), Gilmer (SD83,
Lo82*, UN, WMC), Habersham (JD), Lumpkin (SD83), Murray (GB2, KT), Taylor (SD),
White (GB3, KT).
Uncommon
in mountains and rare in Piedmont in fast clean streams. [May 15 - Jun 23].
Gomphus
(Gomphurus) vastus Walsh, 1862. Cobra Clubtail. State: N&W (1955*), Westfall (1974),
NW&M (2000), Donnelly (2004b).
Appling (GB1, GB2), Burke
(Cr55b, GB2), Cobb (GB), Emanuel (GB1), Meriwether (MV3), Monroe (GB3, MV3),
Newton (MV3), Rockdale (FS2), Toombs (GB2), Treutlen (GB1, GB2), Upson (GB1).
Uncommon
to locally common along rivers and streams south of the mountains. [Apr 11 - Jun 30].
Gomphus
(Gomphus) australis Needham, 1897. Clearlake Clubtail. * State: Donnelly (2004b).
Taylor (SD), Telfair (MD4,
GB4)
Very
rare at sand-bottomed lakes in the coastal plain. The Taylor County record is: Patsiligia
Reservoir at Rt 19, 4 mi. N of Butler, 2 May 1987, 4 males. Collected and
determined by Sid Dunkle. The Telfair
County record is: Cedarpark, 8.5 ml SSE
of McRae, “Brewer’s Pond”, Photos and capture / release, 23 Apr & 28 Apr
2004, males, Marion Dobbs & Giff Beaton. Cross (1955a, 1956) reports it
from Lake Bedford and Dog Lake in Leon County FL. Dunkle(1989) reports it at scattered locations across northern
Florida, so it should be found at more locations across extreme southern
Georgia. [Apr 23 - May 2].
Gomphus (Gomphus) diminutus Needham, 1950. Diminutive Clubtail. State: NW&M (2000), Donnelly (2004b).
Richmond
( WM98*, GBi03).
The
three cited records all refer to a single collection on a
small stream in the eastern part of the state. The locality is: Richmond Co., South Prong of Spirit Creek at US Hwy 1, 16 miles N.
of Wrens, about noon, May 5, 1985; collected and determined by Jerrell J.
Daigle. On 13 Apr 2005 Giff Beaton found and photographed two more specimens in
Richmond Co. on Ft Gordon. The female was at Boggy Gut Creek, and the male was
near Brier Creek along an unnamed sandy road. Bick (1983b) reports it from
Moore County, North Carolina and Chesterfield County, South Carolina. [13 Apr - May
5].
Gomphus (Gomphus) exilis Selys, 1854. Lancet Clubtail. State: Montgomery (1947*), Louton (1982), NW&M
(2000),
Donnelly (2004b).
Atkinson (MD4), Baldwin (SP,UN, WMC), Bartow (GB1,
UG), Bibb (SD), Butts (WMC), Carroll (GB2), Chattooga (GDS3, SG3), Clarke (FSCA),
Cobb (GB), Colquitt (MV3), Cook (MV3), Coweta (GB2), Crawford (SD), Dade
(GDS3), Dawson (MW9), Early (GB4), Emanuel (GB4), Fannin (GB3, GB4), Floyd
(MD3), Forsyth (RT), Gilmer (GB2, GB3, WMC), Gwinnett (GB), Habersham (SD,
FSCA, JD), Harris (SD, GB3, MV3), Heard (GB4), Houston (UG), Jackson (FSCA,
UG), Jones (GB3, MV3, WMC), Lanier (MD4), Lumpkin (SD, MV3), Meriwether (MV3),
Monroe (SD, MV3), Morgan (FSCA, MV3), Murray (SD, GB2, KT98, WMC), Putnam (UG),
Richmond (SD, UN), Rockdale (FS2),
Stewart (MV3), Talbot (SD, GB1), Taliaferro (FSCA, UG), Taylor (SD, GB3),
Telfair (MD4), Union (FSCA, KT), Walker (GDS3), White (SD, FSCA, KT), Wilkinson
(DP).
Common
throughout the state except for the extreme southeast at ponds and lakes; rare
along streams. [Mar 17 - Jul 4].
Gomphus (Gomphus) lividus Selys, 1854. Ashy Clubtail. State: N&W (1955*), NW&M (2000), Donnelly (2004b).
Baldwin (SP), Bartow (GB2, GB), Bleckley (MD4),
Chattooga (GDS3, SR), Cherokee (GB1), Clarke (UG), Cobb (GB), Coweta (GB2, UN),
Crawford (SD), Crawford / Bibb (SD), Dade (GDS3), Dawson (WMC), Floyd (MD3),
Franklin (WMC), Greene (FSCA), Habersham (SD, FSCA, JD), Harris (SD), Jones
(MV3), Lumpkin (MV3), Marion (SD), Meriwether (MV3), Monroe (SD, MV3), Murray
(KT98, WMC), Rabun (UN), Richmond (FSCA), Rockdale (FS2), Seminole (SD),
Stewart (MV3), Talbot (SD), Taylor / Schley (SD), Towns (GB2), Treutlen (GB2),
Union (KT), Walker (SD, GDS3, SR), White (GB4), Whitfield (GB2, Lo82, UN).
Common
above the fall line, uncommon below in the northern coastal plain; occurs at
all sizes of streams and rivers and rarely at lakes and ponds. [Mar 17 - Jun 2].
Gomphus (Gomphus) minutus Rambur, 1842. Cypress Clubtail. State: Selys (1854b*, 1858), Hagen (1861, 1863, 1874,
1875), Kirby (1890), Banks (1892), Calvert (1893), Muttkowski
(1910), N&H (1929), Byers (1930),
N&W (1955), NW&M (2000), Donnelly (2004b).
Ben Hill (WMC), Brantley (BB, DC89b), Bryan (UN),
Burke (Lo82, UN), Calhoun (MV3), Candler (Lo82, UN), Charlton (SD, KT, OO,
WMC), Clinch (TD, WMC), Echols (WMC), Emanuel (SD, FSCA, GB1, KT), Evans (KT,
WMC), Gwinnett (PA), Houston (UG), Jasper (MV3), Laurens (GB1), Long (GB3, KT),
Lowndes (WMC), Pierce (SD), Tattnall (JD, KT, SK, WM95a, WMC ), Telfair (SD,
GB4, WMC), Toombs (KT, SK, WMC), Twiggs (GB4), Wayne (DS, DP), Whitfield
(Lo82).
Fairly
common below the fall line; uncommon to rare above at slower rivers, lakes and
ponds. [Mar 26 - Jun 18].
Gomphus (Gomphus) quadricolor Walsh, 1862.Rapids Clubtail. State: NW&M (2000*), Donnelly (2004b).
Murray
(SK)
Known from only one record in far north Georgia on a clean fast stream:
Murray Co., Conasauga R,
Alaculsy Valley, 5 ml E of Cisco on old hwy 2, 1000 ft., 1 female, 16 May 1998,
Coll. S. Krotzer. It should occur
across the northern ten percent of the state. [Only date recorded: May 16].
Gomphus (Hylogomphus) adelphus Selys, 1854.
Mustached Clubtail. State: Donnelly (2004b).
Murray: ( WM03*).
The single female specimen was collected and photographed by Giff Beaton in northwestern Georgia. The record is: Murray Co., Lake Conasauga Songbird Mgmt. Area, shallow beaver pond and feeder streams, N34° 51.73’, W 84° 39.83’, 9 Jun 2002, Coll. G. Beaton. This is the southern-most record for this species. [Only date recorded: Jun 9].
Gomphus
(Hylogomphus) apomyius Donnelly, 1966. Banner Clubtail. State: NW&M (2000),
Donnelly (2004b).
Bryan / Evans (GB), Crawford
/ Bibb (SD), Meriwether (MV3), Monroe (SD, GB3, MV3), Putnam (MV3), Richmond
(WM98*), Talbot (SD, JD, WM98*), Tattnall (DS), Upson (SD).
Rare
or local at riffles on rivers in the central part of the state; possibly
overlooked due to early flight season. [Apr 7 - May 12].
Gomphus (Hylogomphus) geminatus Carle, 1979. Twin-striped
Clubtail. State: NW&M
(2000), Donnelly (2004b).
Decatur (Ca79*); Taylor (JD, SK, GB3).
Very rare in extreme southwest Georgia, plus a cluster of records in the
western fall line sandhills. It is interesting to note that the Taylor County records are
possibly an isolated population. Additional collecting is needed to determine
if this is indeed a disjunct population. In Florida Dunkle (1992) reports it only west of Tallahassee.
[May 25 - May 30].
Gomphus (Hylogomphus) parvidens Currie, 1917. Piedmont Clubtail. State: Bick (1983b), Dunkle (2000), NW&M (2000)
as G. p. carolinus, Donnelly (2004b).
Fannin (SD), Gilmer (Lo82*, UN), Lumpkin (SD), Rabun
(SK3), Richmond (SD83, KT, JD, UN), Taylor / Schley (SD), White (GB4).
Rare
in the northern half of the state to just below the fall line at rivers and
streams. [May 12 - Jun 6].
Hagenius brevistylus Selys, 1854. Dragonhunter. State: N&W (1955*), NW&M (2000), Donnelly (2004b).
Baker (GB1), Bryan (DS), Burke (Wi34), Butts (MD4),
Carroll (GB3), Catoosa (Lo82), Cherokee (GB2), Dade (GB2, Lo82, UN), DeKalb
(UG), Evans (DS), Floyd (GB2), Gilmer (GB3), Gordon (SD), Habersham (SD),
Hancock / Warren (SD), Haralson (GB2), Murray (GB2, MD4), Oglethorpe (GBi) Polk
(GB2), Rabun (SK3), Rockdale (FS2), Tattnall (DS), Union (GB3).
Fairly
common above the fall line and in the eastern coastal plain except absent from
the southernmost part, on streams and rivers. [Jun 6 - Sep 28].
Lanthus vernalis Carle, 1980. Southern Pygmy Clubtail. State: NW&M (2000), Donnelly (2004b).
Fannin (GB5), Gilmer (Lo82), Lumpkin (WMC), Murray
(SR), Union (SD, GB4), White (Ca80*; SD, JD, WMC).
Rare
to uncommon in the Blue Ridge region along or near trickles and streams. It reaches its southern limit in Georgia, mostly on
the eastern side of the Appalachians. [May 17 - May 28].
Ophiogomphus edmundo Needham, 1950. Edmund's
Snaketail. State: NW&M
(2000),
Donnelly (2004b).
Murray (WM98*, KT98, GBi03, GB4), White
(WM98*, KT98, GBi03).
Very rare in the Blue Ridge region on fast clean forested streams with
exposed rocks. It was discovered in North Carolina in the early 1900s and was reported by Bick (1983b) as “probably extinct.” Rediscovered in North Carolina in 1994, it
is rare in collections, since the adults probably forage in treetops and
typically only drop down momentarily to mate over small rock strewn riffle
streams. In 1998 the senior author stood for some time in the shade, watching a
rock at the head of riffles in a partial sunny situation and waiting for males
of edmundo to swoop down out of the treetops and land momentarily
on the rock. With net poised to strike, he caught only a few. In 2004 the
junior author found several males at a different location perching on the same
rocks for over an hour. [Apr 24 -
May 25].
Ophiogomphus incurvatus incurvatus Carle, 1982. Appalachian Snaketail. State: Bick (1983b), Donnelly (1987), NW&M (2000), Donnelly (2004b).
Talbot (SD), White (Ca82*).
Very
rare above the fall line in clean streams; known only from two records. On 6 Apr 2005 Giff Beaton collected an adult male
near an unnamed stream in Early Co. that shows characters of both O. incurvatus incurvatus and O incurvatus alleghaniensis. [No date
information].
Ophiogomphus mainensis Packard, 1863. Maine Snaketail. State: Donnelly (2004b).
Murray (WM98*, KT98), Rabun (SK3).
Donnelly (1987) also reports it from Oconee
Co SC, adjacent to Rabun County. It is very rare in
the Blue Ridge region in clean fast streams, known only from these two records.
It reaches its southern limit in Georgia. [Only date recorded: May 17].
Progomphus obscurus (Rambur, 1842). Common Sanddragon. State: Hagen (1874*, 1875) and Selys (1878a) as P.
borealis, Banks (1892), Calvert (1901, 1906), Kennedy (1917), N&W
(1955), NW&M (2000), Donnelly (2004b).
Baldwin (SP), Banks (UN), Berrien (GB3), Bryan (UN),
Bulloch (GB2), Burke (GB2), Butts (WMC), Chattooga (MD4), Cherokee (Lo82), Clay
(GB2), Coffee (DP), Columbia (UN), Cook (MD3), Coweta (UG), Dawson (GB4),
DeKalb (UG, UN), Early (MD4), Echols (MD4), Elbert (GB), Emanuel (GB1, GB4),
Evans (DS), Fayette (KT), Franklin (WMC), Fulton (GWQ71), Grady (GB1, UG), Gwinnett (UG), Habersham (SD), Hall
(MW9), Hancock / Warren (SD), Haralson (MD4), Houston (SD, GB3, JD), Jackson
(UN), Johnson (By31, By39), Lowndes (UG), Macon (GB2), McDuffie / Warren (SD),
Monroe (MD3), Montgomery (GB2), Oconee (MD3), Oglethorpe (MD3), Paulding (GB2),
Pickens (GB), Pierce (JF4), Putnam (GB2), Richmond (Lo82), Rockdale (FS2),
Talbot (SD), Tattnall (DS, GB1), Taylor (GB3), Telfair (DP), Thomas (UG), Tift
(GB3), Treutlen (GB1), Troup (GWQ71), Twiggs (GB3), Ware (RB,
UN), Wayne (By31, By39), White (GB3).
Common
statewide in sandy rivers and streams, more numerous below the fall line. [May 9 - Aug 23].
Stylogomphus albistylus (Hagen in Selys, 1878). Eastern Least
Clubtail. State: NW&M (2000), Donnelly (2004b).
Chattooga (MD4), Cherokee (Lo82*), Coweta (GWQ71), Floyd (MD3), Gilmer (Lo82*), Greene (Lo82*), Gwinnett (GWQ71), Habersham (SD), Murray (Lo82*, KT, KT98), Pickens
(GB4), Rabun (SK3), Towns (UG), Walker (SD, GDS3), White (GB3).
Uncommon
in the northern half of the Piedmont and mountain regions in riffle areas of
clean streams of almost any size. [May 23
- Jul 17].
Stylurus amnicola (Walsh, 1862). Riverine Clubtail. State: NW&M (2000), Donnelly (2004b).
Houston (SD83*).
Known
only from one location on a large sandy river near the fall line. It has not been located there in recent
years in spite of intensive searching. [Jun
21 - Jul 3].
Stylurus ivae (Williamson, 1932) Shining Clubtail. State: N&W (1955), NW&M (2000), Donnelly (2004b).
Brantley (DP), Bryan (DS), Burke (Wi32*, Wi34), Lee
(Wi32*), Toombs / Tattnall (Lo82).
The type locality is Brier Creek in Burke County,
which is described as a “20 - 30 foot wide flowing stream mostly 2 - 3 feet
deep with some 8 - 10 foot holes. It meanders through a swamp forest in a bed
of sand and muck, full of logs and tree tops.” It is uncommon to rare
on smaller sandy rivers and streams below the fall line, and it is probably
overlooked due to its late flight season,. [Sep
3 - Oct 12].
Stylurus laurae (Williamson, 1932). Laura’s Clubtail. State: Montgomery (1947), N&W (1955), NW&M
(2000),
Donnelly (2004b).
Dade (Lo82, UN), Gilmer (Lo82), White (Wi32*, Wi34).
Known
only from three records above the fall line in clean streams. However, Westfall (1953) reports it from Gadsden
County, Florida, so it should be found in the south also. Williamson (1932) reported it at “Baggs
Creek, which flows in rock, sand and mud, 10 - 15 feet wide……swift and shallow
with frequent low rapids. The 2 males were taken on a short stretch where
vegetation afforded some shade to the creek.” [Only date recorded: Sep 25].
Stylurus notatus (Rambur, 1842). Elusive Clubtail. State: Louton (1982), NW&M (2000), Donnelly (2004b).
Floyd (Ko60*).
Only
one record from the northwest corner of Georgia, and at the southern limit of
the species’ range. It occurs at rivers
and lakes. [Only date recorded: Jul 13].
Stylurus plagiatus (Selys, 1854). Russett-tipped Clubtail. State: N&H (1929*), N&W (1955), Louton
(1982), NW&M (2000), Donnelly (2004b).
Appling (DS), Baldwin (SP), Bryan (DS), Burke (Wi34),
Carroll (GB3), Charlton (SD), Cherokee (GB2, UN), Clarke (UG), Clay (SD),
DeKalb (UG), Dougherty (RW67), Early (SD, GB4), Floyd (By31, GB2), Fulton (UN),
Greene (MD3), Heard (GB2), Houston (SD), Lee (GB2), Macon (GB2), Meriwether
(SD), Tattnall (DS, GB1), Telfair (MD3).
Fairly
common on faster stretches of rivers and streams statewide; more common below
the fall line; rare in the mountains. [Jul 10 - Nov 5].
Stylurus scudderi (Selys, 1873). Zebra Clubtail. State: N&W 1955), Louton (1982), NW&M (2000), Donnelly (2004b).
White (Wi23b*, Wi34).
Known
only from one record in the extreme northeast.
It occurs in clean streams. This is the southernmost record for this
widespread northern and midwestern species.
[Only date recorded: Sept 25].
Stylurus spiniceps (Walsh, 1862). Arrow Clubtail. State: NW&M (2000), Donnelly (2004b).
Cherokee (Lo82), Rabun (Ko60*), Union (SD).
Known
only from three records primarily in the northeast section of the state but
including one from the Piedmont, on sandy rivers. [Only date recorded: Sep 18].
Cordulegaster
bilineata (Carle, 1983). (Zoraena of some authors). Brown
Spiketail. State:
N&W (1955) as diastatops), NW&M (2000), Donnelly (2004b).
Baldwin (SP), Burke (DS,
GB3), Dawson (Lo82 as diastatops), Habersham (Ca83*, SD, KT), Murray
(Ca83*, KT, SK), Richmond (Cr55b as C.
diastatops, KP87, Ca83*), Union (Ca83*, SD, KT), Walker (GDS3), White (WMC)
Uncommon
to rare and local above the fall line, except uncommon in the northeast section
of the state, at or near sandy seeps and small forest streams. [Apr 4 - Jun 17].
Cordulegaster
erronea Hagen in Selys, 1878. (Kalyptogaster of some
authors). Tiger Spiketail. State: Carle (1983), NW&M (2000), Donnelly (2004b).
Chattooga (MD4), Floyd
(MD4), Gilmer (Lo82), Habersham (GBi78*), Lumpkin (SD), Murray (GB3, MD4),
Rabun (SD), Towns (SD00, KT), Union (SD, JD, UN), Walker (MD4), White (SD, JD,
KT).
Uncommon
across the far northern section of the state at large rocky seeps and small
forest streams. [Jun
26 - Sep 4].
Cordulegaster
maculata Selys,1854. (Pangaeagaster of some authors).
Twin-spotted Spiketail.
State: Selys (1854b*, 1858, 1878a), Hagen (1861, 1863, 1874, 1875), Banks
(1892), Calvert (1893), N&W(1955), Carle (1983), NW&M (2000), Donnelly
(2004b).
Bartow (GB2), Burke (DS),
Chattooga (MD4), Cherokee (Lo82), Clarke (UG, WMC), Cobb (GB, UN), Early (GB4),
Emanuel (DS), Evans (DS), Fannin (GB2), Gilmer (Lo82), Greene (Lo82), Habersham
(SD), Hall (FSCA), Murray (SD, FSCA, SK, Lo82, WMC), Paulding (WMC), Rabun
(SK3), Rockdale (FS2), Tattnall (SK), Union (GB1), White (WMC), Whitfield
(Lo82, UN).
Fairly
common above the fall line, uncommon in the eastern coastal plain, in streams
ranging from tiny to fairly large. [Mar 17 - Jun 20].
Cordulegaster obliqua
fasciata Rambur, 1842. Southern Arrowhead Spiketail. State: Selys (1854b*, 1858,
1878a as fasciata), Hagen (1861, 1863, 1874, 1875), Banks (1892), Byers
(1930), Carle (1983), N&W (1955), NW&M (2000) as C. o. fasciata and C. o. obliqua,
Donnelly (2004b).
Coffee (DS), Emanuel (DS), Grady (DS2), Long (DS),
Murray (JD, WMC), Sumter (RB), Talbot (SD).
Rare statewide at seeps and
small forest streams. [May 16 - Jul 24].
Cordulegaster obliqua obliqua (Say, 1839). Northern Arrowhead
Spiketail.
State: NW&M, 2000*).
Floyd (MD4), Murray (FSCA).
The single published record is based on a
female in the FSCA., collected in Murray County: Eton, CCC Camp Rd @ Tom Terry Road, N 34° 48.9’ W 84°44.3’,
female, 16 May 1998, Coll. Bill Mauffray.
The Floyd County record is:
Berry College, 4 Jul 2004, Photo by Nelson Dobbs. The dot in Donnelly
(2004b) for C. obliqua at Murray County
refers to both subspecies. This subspecies is rare in extreme northern Georgia.
[May 16 - Jul 4].
Cordulegaster sayi
Selys, 1854. Say's Spiketail. State: Selys (1854b*, 1858, 1869), Hagen (1861), Hagen (1863, 1874,
1875) “near Ogeechee River” Banks (1892), Muttkowski (1910), N&H (1929),
N&W (1955), Dunkle & Westfall (1982), Louton (1982), Carle (1983),
Dunkle (1989, 1995, 2000), NW&M (2000), Donnelly (2004b).
Candler (DS), Coffee (GBi03, DS), Emanuel (GBi03, DS), Evans (GBi03, DS, GB1, JD, KT,
WM95, WMC, UN), Liberty (GBi03, DS), Tattnall (GBi03, JD, KT, SK, WM95, 1995b,
WMC), Thomas (Ca04, GBi83b, WM95), Toombs (GBi03, DS), Wayne (GBi03, DS).
The
Thomas County specimen is the type according to Bick (1983b). Uncommon and local in turkey oak seep systems of
the eastern coastal plain. [Mar 8 - Apr 4].
Didymops transversa
(Say, 1839). Stream Cruiser. State: Hagen (1863*, 1874, 1875), Selys (1871), Banks
(1892), Calvert (1893), Muttkowski (1910), N&H (1929), Byers (1930),
N&W (1955), NW&M (2000), Donnelly (2004b).
Baldwin (SP),
Bartow (GB2, UG), Bleckley (MD4), Brooks (SD), Charlton (WMC), Chattooga (SR),
Cobb (GB), Cook (SD), Coweta (FSCA), Crawford (SD), Decatur (UG), DeKalb (UG),
Effingham (SD, FSCA), Emanuel (KT), Evans (WMC), Fannin (MV3), Habersham (SD,
UG), Harris (SD, MV3), Jackson (UG), Lee (UG, UN), Long (GB3), Lumpkin (SD,
MV3), Meriwether (MV3, UG), Monroe (GB3, MV3), Morgan (MV3), Murray (SD, KT, Lo82, SR), Muscogee / Chattahoochee
(GWQ71), Newton (MV3), Rabun (UG),
Rockdale (FS2, UG), Tattnall (SK, WMC), Taylor / Upson (SD), Thomas (UG),
Toombs (KT, WMC), Towns (GB2), Union (KT).
Fairly common above the fall line, uncommon below; at streams and small
rivers. [Mar 13 - Jun 12].
Macromia
alleghaniensis Williamson, 1909. Allegheny River Cruiser. State: N&W(1955*),
NW&M (2000), Donnelly (2004b).
Dade (Lo82, UN), Troup (GWQ71).
The Dade records are based
on a reared male collected at Lookout Creek and CR 201, 10 May 1980, emerged 14
June 1980; and two females from Lookout Creek, one mile downstream from Easley
Cemetery 18 Aug 1980. It is very rare at
rivers in extreme western Georgia; but since Donnelly (1989) reports it from
Caldwell County, North Carolina and also from northwest South Carolina
(Donnelly, 2004b), it should be found farther east in northern Georgia. [Jun 14
- Aug 18].
Macromia illinoiensis georgina (Selys, 1878). (Macromia georgina of some authors).
Georgia River Cruiser. State: Selys (1878b*), Banks (1892), Calvert (1893),
Martin (1906), Muttkowski (1910), Needham & Heywood (1929), Montgomery
(1945), Needham & Westfall (1955), NW&M (2000), Donnelly (2004b).
Appling (GB2),
Baldwin (SD, SP), Burke (CC89c, DT94, UN), Clay (SD),Clinch (WMC), Dade (DT94,
Lo82), Decatur (UG), Early (SK4), Floyd (GB), Haralson (GB2), Houston (SD,
GB3), Laurens (By31), Lee (DT94), Madison (UG), Meriwether (DT94, SD), Monroe
(GB3), Montgomery (GB1), Murray (MD4), Muscogee / Chattahoochee (GWQ71), Pierce
(DT94), Rockdale (FS2), Talbot / Upson
(GB2), Telfair (MD3), Twiggs (GB3), Wilkes (UG), Wilkinson (SD).
Fairly common statewide on larger streams and rivers. [May 11 - Sep 22].
Macromia
margarita Westfall, 1947. Mountain River Cruiser. State: NW&M (2000), Donnelly (2004b).
Lumpkin (GBi03, Ko60*).
Very
rare at rivers in extreme northeast Georgia; known from only one record. Bick
(2003) and Donnelly (2004b) summarize its distribution ranging from northwest
Alabama northward into Tennessee and eastward into North and South Carolina.
Based on this range it should occur in the upper fifteen percent of the state. [Only date recorded: Jul 2].
Macromia
taeniolata Rambur, 1842. Royal River Cruiser. State: Hagen (1863*, 1874, 1875), Banks
(1892), Calvert (1893), Martin (1906), N&W (1955), Louton (1982), NW&M
(2000), Donnelly (2004b).
Appling (CC89c), Bartow
(CC89c), Burke (By31, Wi34), Decatur (UG), Emanuel (GB4), Houston, (CC89c),
Laurens (CC89c), Lee (Ro24, CC89c), Long (GB3), Lowndes (CC89c), Sumter (UG),
Tattnall (GB, GB1), Taylor (CC89c), Telfair (DP), Wilkes (CC89c), Wilkinson
(DP).
Fairly
common at rivers below the fall line, uncommon above. [May 15 - Sep 21].
Corduliidae -
Corduliinae
Epitheca
(Epicordulia) princeps (Hagen, 1861). Prince Baskettail (includes all
references to E. regina ) State: Hagen (1861*, 1863,
1874, 1875), Selys (1871), Kirby (1890), Calvert (1893), Martin (1906), Muttkowski (1910), N&H (1929), Byers (1930),
N&W (1955), NW&M (2000), Donnelly
(2004b).
Baldwin (SD, SP), Bibb (UG),
Brooks (GB1), Carroll (GB2), Catoosa (GB), Cobb (GB), Cook (GB1), Decatur (UG),
Dougherty (RW67), Douglas (GB), Early (SK4), Fayette (KT), Floyd (GB1), Gordon
(GB2), Habersham (GBi), Hall (MW9), Hancock / Warren (SD), Jasper (GB2),
Laurens (By31, as regina, GB),
Lee (Ro24, RB), Long (GB4), Meriwether (GB1), Monroe (GB3, MD3), Pickens (GB), Polk
(GB), Putnam (GB2), Rabun (GBi) Rockdale (FS2), Stephens (GB2), Sumter (UG),
Taliaferro (JF4), Tattnall (DS), Union (SD),
Upson (GB1), Walker (GDS3).
Common
statewide except uncommon in the southeast at ponds, lakes, and rivers. [May 2 - Sep 24].
Epitheca
(Tetragoneuria) costalis (Selys, 1871). Stripe-winged Baskettail. State: Hagen (1861*, 1874,
1875), Selys (1871), Kirby (1890), Banks (1892), Martin (1906), Muttkowski (1910), NW&M (2000), Donnelly (2004b).
Baldwin (SP), Brantley (BB),
Chattooga (GDS3), Clinch (KT77), Cobb (GB3), Crawford / Bibb (SD), Dade (GDS3),
Long (KT, WMC), Lowndes (MD4), Murray (SK), Tattnall (JD, KT, SK, WM95a),
Walker (GDS3).
Locally
uncommon to rare statewide at ponds and lakes. It may be overlooked. [Mar 26 - Jun 20].
Epitheca
(Tetragoneuria) cynosura (Say, 1839). Common Baskettail. State: Hagen (1863*, 1874)
as Cordulia lateralis, (1875), Banks (1892), Muttkowski
(1915), N&W (1955), NW&M (2000), Donnelly
(2004b).
Bacon (GB1), Baldwin (SP),
Bartow (GB2), Bibb (UG), Brantley (BB), Brooks (UG), Bulloch (DP), Calhoun
(MV3), Carroll (GB2), Charlton (OO), Chatham (UG), Chattooga (MD4, GDS3)),
Cherokee (GB1), Clarke (UG), Clinch (KT), Cobb (GB), Coffee (GB3), Colquitt
(MV3), Columbia (UN), Cook (MV3), Coweta (GWQ71, UG), Dade (GDS3), Dawson
(MW9), DeKalb (UG), Early (GB3, GB4, MV3), Effingham (GB4), Emanuel (KT), Evans
(KT), Floyd (MD4), Fulton (UG), Gilmer (WMC), Gwinnett (PA), Haralson (WMC),
Harris (MV3, UG), Heard (GB4), Houston (UG, UN), Jackson (UG), Johnson (UG), Jones (MV3, WMC), Laurens (GB3), Long (KT,WMC), Lowndes (MD4), Lumpkin (MV3),
McIntosh (MV3), Meriwether (MV3), Monroe (MV3), Morgan (MV3, UG), Murray (KT,
SR, WMC), Newton (MV3), Oconee (UG),
Paulding (GB, GB3), Rabun (GBi), Richmond (UN), Rockdale (FS2), Taliaferro
(UG), Tattnall (KT, SK, MV3, WM95a), Taylor (GB3), Telfair (UN), Toombs (KT),
Union (GB1, KT), Walker (GDS3), Ware (MV3), Wayne (DP), Wheeler (GB1), White
(KT)
Common
statewide at marshes, ponds, and lakes, and uncommon at slow streams and
rivers. [Mar 13
- Jul 4].
Epitheca
(Tetragoneuria) semiaquea (Burmeister, 1839). Mantled Baskettail. State: Hagen (1863, 1873,
1874), Selys (1871), Banks (1892), Muttkowski
(1911a), Davis (1933) as calverti, N&W (1955), NW&M
(2000), Donnelly (2004b).
Charlton (OO, DP), Chatham
(Bu39*, Ca98, Ha61, Ha75), Clinch (TD, UG), Emanuel (KT), Evans (KT, WMC),
Houston (UG), Long (JD, KT, WMC), Richmond (UN), Tattnall (JD, SK, WMC, WM95a);
Thomas (Da33, Mu15), Toombs (KT), Wayne (DP).
It was first published as Libellula semiaquea from the type
locality of “Savannah” Burmeister (1839). Uncommon
to locally common at ponds and lakes in the eastern coastal plain. [Mar
18 - Apr 13].
Epitheca (Tetragoneuria) sepia Gloyd, 1933. Sepia Baskettail. State: NW&M
(2000), Donnelly
(2004b).
Tattnall
(WM95a), Taylor (SK3), Ware (GBi90*), Wayne (DP).
Rare in the coastal plain at ponds, lakes and slow streams and rivers;
mostly active at dusk. It may be overlooked. [May 25 - Jul 8].
Epitheca (Tetragoneuria) spinosa (Hagen in Selys, 1878). Robust Baskettail. State: Selys
(1878b*) Banks (1892), Martin (1906), Muttkowski (1910, 1915), N&H (1929),
N&W (1955), NW&M (2000), Donnelly (2004b).
Bick (1983a) reports a specimen from Tuscaloosa,
Alabama. Dunkle (1989) reports only one record from north Florida in the
panhandle. Kondratieff & Pyott (1987) report it from Savannah River Plant
just across the border in South Carolina. All the Georgia records refer to the
original Selys (1878b) record. Although there are no modern records, its range
according to Donnelly (2004b) would seem to indicate that it should continue to
occur in Georgia. [No date information].
Epitheca (Tetragoneuria) stella Williamson in Muttkowski, 1911. Florida Baskettail. State: Byers
(1930), N&W (1955), NW&M (2000), Donnelly (2004b).
Lee (UG), Wayne
(By31), Thomas (Mu15*).
Rare in southern coastal plain at ponds, and also may be overlooked. [Only date
recorded: Mar 16].
Helocordulia
selysii (Hagen in Selys, 1878). Selys' Sundragon. State: Selys (1878b*),
Kirby (1890), Banks (1892), Martin (1906), Muttkowski
(1910), N&H (1929), N&W (1955), NW&M
(2000), Donnelly (2004b).
Lumpkin (MV3), Monroe (MV3),
Morgan (MV3), Paulding (GB3), Rockdale (FS2), Taylor (GB3).
Rare to uncommon in the Piedmont
and upper Coastal Plain at sandy streams. [Mar 17 - May 1].
Helocordulia
uhleri (Selys, 1871). Uhler's Sundragon. State: NW&M (2000), Donnelly
(2004b).
Bartow (GB2, GB4), Floyd
(MD4), Murray (WM98*, GB4).
Rare
to uncommon in the northwestern portion of the state at clean streams and
rivers. [Apr 6
- May 25].
Neurocordulia
alabamensis Hodges in Needham & Westfall, 1955. Alabama
Shadowdragon. State:
N&W(1955*), Dunkle (1989), NW&M (2000), Donnelly
(2004b).
N&W55 report the type
locality as Alabama, but also list it from Georgia, Florida, and South
Carolina. Louton
(1982) reports it from Aiken County, South Carolina just across the river from
Georgia. Donnelly (2004b) shows scattered records across the Gulf and Atlantic
coastal plains. Because it, along with the other species of this genus, is
active only at dusk, it is uncommon in collections. [No date
information].
Neurocordulia molesta (Walsh, 1863). Smoky Shadowdragon. State: Westfall (1953*) “Savannah
River”, N&W (1955), NW&M (2000), Donnelly (2004b).
Bartow (Lo82),
Jeff Davis / Montgomery (Lo82), Seminole (GWQ71). Long (GB5), Telfair / Jeff
Davis (GB5)
Rare and probably overlooked throughout the state at rivers, it is known
from only five records. Louton (1982) also reports it from a few counties in South
Carolina along the Savannah River. [Exuviae records 15 May - 16 May].
Neurocordulia
obsoleta (Say, 1839) Umber Shadowdragon. State:
NW&M (2000), Donnelly (2004b).
Morgan (MV3). Toombs (WM98*)
The Morgan County record
is: Rutledge, Hard Labor Creek State
Park, Hard Labor Creek, 23 Apr 2003, Coll. by Michael Veit. The Toombs County
record is: Cobb Cr., Hwy. 147, 1 Apr 95, 1 female, Coll. K. J. Tennessen.
Williamson (1903b) reports a larval record from Chattanooga, Tennessee. Based
on its range (Donnelly, 2004b), it probably occurs throughout most of the
state, but it is very rare. [No date
information].
Neurocordulia virginiensis Davis, 1927. Cinnamon Shadowdragon. State (Byers
1937*), N&W (1955), NW&M (2000), Donnelly (2004b).
Dougherty
(RW67), Early (SK).
The Dougherty
County records are nymphs. The Early County record is of two males collected by
Steve Krotzer on 2 May 1999 from the west bank of the Chattahoochee River out
over the river, which is in Georgia (per. comm.). Byers’ (1937) holotype was
from Jackson County, Florida. Louton (1982) reports it from the Conasagua River
in Tennessee and in a couple of counties along the Savannah River near Aiken,
South Carolina. It is widespread, but rare, in clean rivers. [No date
information].
Somatochlora
elongata (Scudder, 1866). Ski-tailed Emerald. State: NW&M (2000), Donnelly (2004b).
White (SD83*).
It
reaches the southern most extreme of its range in northeast Georgia. It is very
rare at slow streams and marshes. [No date information].
Somatochlora
filosa (Hagen, 186l ). Fine-lined Emerald. State: Hagen (1861*, 1863, 1874, 1875), Selys
(1871), Kirby (1890), Banks (1892), Calvert (1893), Martin (1906), Walker
(1925), N&W (1955), NW&M (2000), Donnelly
(2004b).
Brantley (DP), Charlton
(GBi, SD, GB4, UN), Clarke (UG), Clinch (SD).
Uncommon
to rare in Georgia, mostly below the fall line, near rivers but breeding
habitat unknown.
[Sep 8 - Nov 4].
Somatochlora
georgiana Walker, 1925. Coppery Emerald. State: N&H (1929), N&W (1955), NW&M
(2000), Donnelly (2004b).
Lee (Wa25*, JD92b), Telfair
(JD92b, DP).
The type locality is in Lee
County. It is rare in the coastal plain at
small forest streams. [No date information].
Somatochlora
linearis (Hagen, 1861). Mocha Emerald. State: Hagen (1874*), Selys (1878b), Banks (1892),
Calvert (1893), Montgomery (1945), N&W (1955), NW&M (2000), Donnelly (2004b).
Bibb (UG), Burke (By31,
Wi34), Clinch (DP), Dodge (UN), Emanuel (SD), Floyd (By31, GB2), Houston (SD,
GB3), Jefferson (Wi34), Lee (Ro24, Wa25), Long (DS), Polk (GB2), Twiggs (GB3),
Wheeler (SD).
Fairly
common statewide at small forest streams; found in greatest density as the
streams are drying up in late summer. [Jun 13 - Sep 14].
Somatochlora
provocans Calvert, 1903. Treetop Emerald. State: N&W (1955), NW&M (2000), Donnelly (2004b).
Lee (Ro24*, Wa25)
Very
rare in coastal plain. It is known from only one record. It occurs at forest seeps and trickles. [Only recorded date: Jul 7].
Somatochlora
tenebrosa (Say, 1839). Clamp-tipped Emerald. State: NW&M (2000), Donnelly (2004b).
Chattooga (GB3), Cobb (GB3),
Dade (WM98*, UG, UN), Floyd (GB2), Fulton (WM98*, UG), Murray (GB3, GB4), Rabun
(GB3, JD, WM98*), Union (GB3), White (SD).
Fairly
common at forest seeps and streams in the northern half of the Piedmont and
mountains, but should also be found in southwestern Georgia since there are
several records near Tallahassee, Florida (Donnelly, 2004b). [Jun 20 - Sep 10].
Brachymesia
gravida (Calvert, 1890). (Cannacria gravida of various authors)
Four-spotted Pennant.
State: NW&M (2000), Donnelly (2004b).
Baker (GB2), Ben Hill (GB2),
Berrien (GB), Brantley (SD75*), Brooks (GB1), Bulloch (AH1, AH2), Chatham (GB),
Coffee (RB), Colquitt (GB, MD3), Crisp (GB2), Dodge (GB1), Emanuel (GB1), Evans
(GB), Glynn (GB2), Grady (GB1), Irwin (SD75*, KT), Jasper (GB2), Jeff Davis
(GB), Laurens (GB), Lee (RB), Liberty (DS, GB), Lowndes (GB), McIntosh (GB),
Miller (GB), Montgomery (GB1), Pierce (JF4),
Putnam (GB2),
Seminole (GB), Sumter (RB), Talbot / Upson (GB2), Tattnall (DS), Tift (RB),
Toombs (GB1), Wilcox (GB2).
Common
at ponds and lakes in the coastal plain, especially those with cattail edges,
and rare in the lower Piedmont. [May 8 - Nov 11].
Celithemis
amanda (Hagen, 1861). Amanda's Pennant. State: Hagen (1863, 1874, 1890b), Kirby (1890),
Banks (1892), Williamson (1922c), N&H (1929), Byers (1930), N&W (1955),
NW&M(2000), Donnelly (2004b).
Brantley (UG), Bulloch (DP),
Charlton (UN), Chatham (Ha61*), Clinch (UG), Glynn (UN), Irwin (SD), Lanier
(GB4, UG), Laurens (GB1), McDuffie (Wi34) Richmond (UG), Taylor (GB4), Thomas
(UN), Treutlen (MD4).
Uncommon
to rare at ponds and small lakes in the eastern coastal plain. [Mar 29 - Sep 28].
Celithemis bertha Williamson,
1922. (as leonora ) Red-veined
Pennant. State: N&W (1955*) also as leonora, NW&M (2000)
as C. b. bertha and C. b. leonora, Donnelly (2004b).
Appling (GB4),
Baker (GB2), Baldwin (SP), Bulloch (AH2), Coffee (MD4, RB), Cook (MD4), Decatur
(FSCA), Dodge (MD4), Floyd (MD3), Glynn (UN), Grady (GB1), Irwin (FSCA),
Laurens (GB1), Lee (RB), Rabun (GB3),
Stephens (GB2), Tattnall (GB1).
Uncommon to fairly common at shallow ponds and lakes with barely emergent
shoreline vegetation in the coastal plain, and occasionally on rivers and lakes
above the fall line throughout the Piedmont. [May 2 - Oct 17].
Celithemis
elisa (Hagen, 1861). Calico Pennant. State: Hagen (1863*, 1874, 1875, 1890b), Calvert
(1893), Ris (1912), Williamson (1922c), N&W (1955), NW&M (2000),
Donnelly (2004b).
Appling (GB4), Atkinson
(MD4), Baker (GB2, UG), Baldwin (MD4, SP), Bartow (UG), Ben Hill (GB2, WMC),
Bulloch (DP), Butts (MD4), Camden (UG), Carroll (GB), Chattooga (GB2), Clarke
(UG), Clay (GB3), Cobb (GB), Coffee (RB), Cook (UG, UN), Coweta (GB2), Crawford
(UN), Dade (GDS3, GB2), Decatur (UG, UN), Douglas (GB), Echols (SD), Elbert
(GB), Emanuel (GB1), Fannin (GB3), Floyd (MD4), Franklin (GB, UG), Gordon
(MD4), Grady (GB1), Gwinnett (UG), Habersham (GBi), Haralson (WMC, GB2), Harris
(GB1, UG), Irwin (SD, GB3), Jones (MV3), Lamar (GB), Laurens (GB, GB1), Lee
(Ro24, RB), Long (GB3), Lumpkin (SD), McDuffie (Wi34), Meriwether (GB1), Miller
(GB), Mitchell (UG), Montgomery (GB1, GB2), Morgan (GB), Murray (GB2, UG),
Paulding (GB), Peach (GB), Pickens (GB), Pierce (SD), Pike (UG), Polk (GB),
Rabun (GBi, SD, GB3), Richmond (SD, UN), Rockdale (FS2, GB), Stephens (GB2),
Talbot (GB1), Taylor (GB4, MD4), Toombs (GB1), Union (GB3, KT), Upson (UG),
Walker (GDS3, GB2), Wayne (UG, UN), Wheeler (WMC), Wilcox (GB2), Worth (WMC).
Common
statewide at ponds and lakes, often with grassy or vegetated edges. [Apr 3 - Nov 21].
Celthemis
eponina (Drury, 1773). Halloween Pennant. State: Rambur (1842*) as Libellula camilla, Hagen (1861, 1863, 1874, 1875), Ris (1912),
N&W (1955), NW&M (2000), Donnelly (2004b).
Baldwin (SP), Bartow (GB),
Ben Hill (GB2), Berrien (GB, MD4), Brooks (GB1), Bulloch (AH1, AH2), Burke
(GB2, Wi34), Butts (MD4), Camden (UN), Catoosa (GB), Charlton (Br14), Chattooga
(GB2), Clarke (UG), Clayton (GB), Cobb (GB), Coffee (RB), Colquitt (GB, MD4),
Crisp (GBi), Decatur (UG), DeKalb (UG), Douglas (GB), Early (GB4), Evans (GB),
Floyd (GB), Franklin (GB), Fulton (UG), Gwinnett (GB), Haralson (GB2), Houston
(UG), Irwin (SD, RB), Jasper (GB2), Jeff Davis (GB), Lee (Ro24), Lowndes (GB,
UN), Meriwether (GB2), Mitchell (UG, UN), Montgomery (GB1), Paulding (GB2),
Polk (GB), Putnam (UG), Rockdale (FS2), Seminole (GB), Sumter (RB), Talbot
(MD4), Telfair (DP), Tift (RB), Toombs (GB1), Wheeler (DP).
Common
statewide but may be local at marshes, ponds and lakes. [May 29 - Nov 28].
Celithemis
fasciata Kirby, 1889. Banded Pennant. State: Kirby (1890), Hagen (1890b), Banks (1892),
Williamson (1910, 1922c), Ris (1912, 1916), Byers (1930), N&W (1955),
NW&M (2000), Donnelly (2004b).
Atkinson (SD, MD4), Baker
(UG), Baldwin (SP), Bartow (GB1), Bibb (UG), Bulloch (AH1, AH2), Butts (MD4),
Carroll (GB2), Charlton (Br14), Chattooga (GB2), Clarke (UG), Clinch (GBi),
Cobb (GB, UG), Coffee (RB), Crawford (UN), Dade (GB2), DeKalb (UG), Early
(SK4), Elbert (GB), Fannin (GB3), Floyd (GB), Franklin (UG), Glascock (JF3), Gwinnett
(GB, JF), Habersham (GBi, SD), Hall (MW9), Hancock (MD4), Haralson (WMC),
Houston (UG), Irwin (SD), Jasper (GB2), Jeff Davis (GB), Lanier (UG), Laurens
(GB1), Lee (RB), Lowndes (UN), Lumpkin (SD), Macon (SD, UG), McDuffie (Wi34),
McIntosh (GB), Meriwether (JF3), Murray (MD4), Peach (UG), Pickens (GB, GB2),
Putnam (GB2, UG), Rabun (GBi, SD), Richmond (UG, UN), Rockdale (FS2, GB),
Stephens (GB2), Taliaferro (JF4), Taylor (UG), Telfair (MD4),
Toombs (GB1), Union (SD), Upson (UG), Walker (GDS3, GB2), Wheeler (DP).
Fairly
common statewide, may be locally common, at ponds and lakes. [May 15 - Nov 5].
Celithemis
ornata (Rambur, 1842). Faded Pennant. State: Hagen (1874*), Ris (1912), Williamson
(1922c), Byers (1930), N&W (1955), NW&M (2000), Donnelly (2004b).
Baker (UG), Ben Hill (WMC),
Bibb (UG), Camden (GB), Charlton (SD, UG), Crawford (UG), Echols (WMC), Evans
(WMC), Glynn (UG), Laurens (GB4), Lee (Ro24, UG), Lowndes (WMC), McDuffie
(Wi34), Pierce (SD, FSCA), Richmond (SD), Taylor (UG), Telfair (MD4), Wheeler
(DP).
Uncommon
in the coastal plain at ponds and lakes with emergent shoreline vegetation. [Mar 25 - Nov 5].
Celithemis
verna Pritchard, 1935. Double-ringed Pennant. State: N&W (1955), NW&M (2000),
Donnelly (2004b).
Baker (UG), Cook (Pr35*), DeKalb
(UN), Evans (UN), Lee (Pr35*), Monroe (GB3), Taylor (GB3), Telfair (MD4),
Walker (GDS3), Wheeler (DP, UN), White (SD).
Rare
statewide at ponds and lakes with emergent shoreline vegetation. [Apr 23 - Aug 6].
Dythemis
velox Hagen, 1861. Swift Setwing. State: NW&M (2000), Donnelly (2004b).
Baldwin (SP), Bartow (MD4),
Butts (MD4), Carroll (GB), Catoosa (GB), Chattooga (GB2), Cherokee (GB2),
Clarke (UG), Clayton (WMC), Early (SK4), Floyd (GB, GB1), Forsyth (RT),
Franklin (MD3), Fulton (UG), Gordon (MD4), Gwinnett (GB, JF), Habersham (GBi,
SD), Hall (MW9), Harris (SD), Lee (Ko60*), Meriwether (SD, GB1), Monroe (GB3),
Paulding (GB2), Polk (GB), Rockdale (FS2, GB), Stephens (GBi), Sumter (MD4,
RB), Whitfield (MD3), Wilkinson (SD).
Fairly
common to uncommon above the fall line and uncommon to rare in the western
coastal plain at streams, rivers, ponds, and lakes. [May 20 - Oct 16].
Erythemis
simplicicollis (Say, 1839). Eastern Pondhawk. State: Hagen (1863, 1874, 1875), Ris
(1911), N&W (1955), NW&M (2000), Donnelly (2004b).
Appling (GB1), Atkinson
(WMC), Baker (MV3, UG, UN), Baldwin (SP), Bartow (GB, GB2, UG), Ben Hill (UG,
WMC), Berrien (GB, GB3), Bibb (GB, UG), Brantley (UG), Brooks (UG), Bryan (GB,
UN), Bulloch (AH1), Burke (UN, Wi34), Butts (MD4), Calhoun (MV3, UG), Carroll
(GB), Catoosa (GB), Charlton (OO, PA, UG, UN), Chatham (GB, Ha61*), Chattooga
(GDS3, GB, GB2, SG3), Cherokee (GB), Clarke (UG), Clay (GB2), Clayton (GB),
Clinch (GBi, WMC), Cobb (GB, UG), Coffee (RB), Colquitt (GB, MD4), Coweta (GB),
Crawford (SD, UN), Crisp (GBi), Dade (GDS3, GB2), Dawson (JF), Decatur (GBi,
UN), DeKalb (UG, UN), Dodge (GB1), Dooly (GB2), Dougherty (UG), Douglas (GB),
Early (GB2, SK4, UG), Echols (WMC), Effingham (GB2, UG), Elbert (GB), Emanuel
(GB1, GB4), Evans (GB), Floyd (GB), Forsyth (GB, JF), Franklin (GB, UG, WMC),
Fulton (UG, HS4), Gilmer (GB3), Glynn (GB, UG, UN), Gordon (UG), Grady (GB1),
Greene (GB), Gwinnett (By31, JF, PA), Habersham (GBi, SD), Hall (MW9), Haralson
(WMC, GB2), Heard (GB2), Henry (UG), Houston (UG), Irwin (SD, GB3, RB, UG),
Jasper (GB2), Jeff Davis (GB), Jefferson (GB2), Jenkins (UG), Johnson (SD),
Jones (WMC), Lamar (GB, UG), Lanier (UG), Laurens (By31, GB, PA), Lee (Ro24,
RB), Liberty (UG), Lowndes (UG, UN, WMC), Macon (GB2), McDuffie (Wi34), McIntosh
(GB, MV3, UG), Meriwether (GB1), Miller (GB), Mitchell (UG, UN), Monroe (GB),
Montgomery (GB1, GB2), Morgan (MV3), Murray (GB, KT, UN), Newton (MV3),
Paulding (GB), Peach (UG), Pierce (SD), Polk (GB), Pulaski (WMC), Putnam (GB2,
UG), Rabun (GBi), Randolph (JF4), Richmond (UG, UN), Rockdale
(FS2, GB, JF), Seminole (GB), Stephens (UN), Stewart (GB, MV3), Sumter (RB,
UG), Taliaferro (UG), Tattnall (WMC), Taylor (SD, UG), Telfair (UG, WMC),
Thomas (UG), Tift (GB3, RB), Toombs (GB1), Treutlen (GB1), Turner (WMC), Twiggs
(GB3), Upson (UG), Walker (GDS3), Ware (MV3, RB, UG, UN), Washington (UG),
Wayne (OO, UG, UN), Wheeler (WMC), White (DP), Whitfield (GB2), Wilcox (GB2),
Wilkinson (DP), Worth (WMC).
Very
common statewide at marshes, ponds and lakes, especially those with lily pads. [Mar 5 - Nov 21].
Erythrodiplax
berenice berenice (Drury, 1770). Seaside Dragonlet. State: Ris (1911*), N&W
(1955), NW&M (2000), Donnelly (2004b).
Bryan (UN), Camden (SD, GB),
Charlton (Br14), Chatham (GB), Glynn (Ca12, SD, SH30, GB1, UG), McIntosh (UG).
Uncommon
along the immediate coast at brackish or salt marshes and wet grassy areas. [Jun 14 - Oct 6].
Erythrodiplax
minuscula (Rambur, 1842). Little Blue Dragonet. State: Hagen (1861*, 1863, 1874, 1875), Banks
(1892), Williamson (1899), Calvert (1906), Ris (1911), Muttkowski
(1910), N&H (1929), N&W(1955), NW&M (2000), Paulson (2003),
Donnelly (2004b).
Appling (GB4, UG), Atkinson
(DP), Bacon (DB42), Baker (UG), Baldwin (SP), Ben Hill (GB2), Berrien (FS2, GB1),
Brantley (BB, UG), Brooks (GB1), Bryan (UN), Burke (Wi34, DB42), Camden (UN),
Charlton (Br14, OO, PA, UG, WMC), Chatham (GB), Chattooga (MD4), Clarke (UG),
Clinch (GB3, UG), Coffee (DP), Cook (MD4, UN), Crawford (UG, UN), Decatur
(DB42, UN), Dougherty (UG), Early (GB3, SK4, UN), Echols (GB3, WMC), Effingham
(UN), Emanuel (GB1, GB4), Glynn (UG), Grady (GB1), Gwinnett (GB, JF), Irwin
(RB), Jefferson (DB42, By31), Jeff Davis (GB, UG), Lanier (GB3, UG),
Laurens (GB1, GB2, WMC), Lee (DB42,
Ro24, RB3, UG, UN), Long (DS, GB3, WMC), Lowndes (DB42, UG, UN, WMC), McDuffie
(UN, Wi34, DB42), McIntosh (GB, GB4, MV3), Mitchell (UG, UN), Montgomery (GB1),
Rabun (GB3), Richmond (UG, UN), Rockdale (FS2, GB, UG), Schley (UG), Screven
(DP), Seminole (GB), Stephens (UN), Sumter (MD4), Tattnall (DS, WMC), Taylor
(UG, UN), Telfair (UG, UN), Tift (MD4), Toombs (GB1, WMC), Treutlen (MD4),
Twiggs (GB3), Ware (DB42), Wayne (DB42, By31, OO, UG, UN), Wheeler (DP), Wilcox
(GB2), Wilkinson (By31, DB42), Worth (WMC).
Prior to Paulson (2003),
this species was called Erythrodiplax connata minuscula. It is uncommon above the fall line, very common
below. It is found at marshes, ponds,
lakes, and slow rivers. [Mar 26 - Dec 8].
Ladona
deplanata Rambur, 1842. (Libellula of some authors). Blue Corporal.
State: Hagen (1861*,
1874, 1875), Banks (1892), Calvert (1893), Ris (1910), N&H (1929),
N&W(1955), Cuyler (1989b), NW&M (2000), Donnelly (2004b).
Baker (MV3, UG), Baldwin
(MD4, SP), Bartow (UG, GB2, UN), Berrien (GB1), Bibb (UG), Brantley (BB),
Brooks (SD), Bulloch (AH3), Burke (GB3), Butts (WMC), Charlton (FSCA, GB2, OO,
UG), Cherokee (GB1), Clarke (UG), Clinch (UG), Cobb (GB), Colquitt (MV3), Cook
(GB1, MV3), Crawford (UG), Crawford / Bibb (SD), Dade (GDS3), Early (GB3, GB4),
Effingham (FSCA) Evans (WMC), Floyd (MD3), Fulton (UG), Gwinnett (GB, PA),
Habersham (SD), Harris (GB3), Heard (GB4), Houston (UG, UN), Jasper (MV3),
Jones (MV3, WMC), Laurens (GB2), Liberty (UG), Long (GB3, WMC), Lowndes (MD4),
Lumpkin (MV3), McIntosh (UG), Mitchell (UG), Montgomery (GB1), Morgan (MV3),
Murray (GB2, KT), Paulding (GB3),
Pulaski (MD4), Rockdale (FS2), Stewart (MV3), Tattnall (GB4), Taylor (GB3,
GB4), Telfair (GB4, MD4), Toombs (GB1), Walker (GDS3), Walton (UG), Ware (MV3),
Wayne (UG), Wheeler (GB1), White (FSCA, KT).
Common
statewide at ponds and lakes. [Mar 4 - Jun 18].
Libellula
auripennis Burmeister, 1839. Golden-winged Skimmer. Hagen (1861, 1863, 1874,
1875), Ris (1910), N&W (1955), NW&M (2000), Donnelly (2004b).
Appling (GB, GB4), Atkinson
(WMC), Bacon (MW43), Baldwin (SP), Bartow (UN), Berrien (GB), Bibb (UG, UN),
Bryan (UN), Bulloch (AH1), Charlton (FS2, GB, PA, UN, MW43), Chatham (Bu39*,
Ca98, Br14, MW43), Clarke (UG), Clinch (SD, GB3, WMC), Cobb (GB), Crawford
(UN), Dawson (JF), Decatur (UG, MW43), Dodge (MD4), Early (GB4, UN), Echols
(GB3), Evans (GB, UN), Fannin (GB3, UN), Floyd (GB), Glynn (By31, GB, UG,
MW43), Grady (GB1), Gwinnett (PA), Irwin (RB), Jeff Davis (GB), Jefferson
(By31,UN, MW43), Lanier (UG, UN), Laurens (GB1, WMC), Lee (Ro24, RB, UG, MW43),
Lowndes (GB), McDuffie (Wi34, MW43), McIntosh (GB, MV3), Mitchell (UN), Murray
(GB2), Randolph (JF4), Richmond (UN), Rockdale
(FS2), Schley (RB3), Seminole (GB), Sumter (MD4), Tattnall (GB4), Taylor (MD4),
Telfair (MD4), Tift (RB), Treutlen (MD4), Walker (GDS3), Ware (RB, UG), Wayne
(UN).
Common
below the fall line, uncommon above, at marshes, ponds, and lakes. [Apr 19 - Oct 23].
Libellula
axilena Westwood, 1837. Bar-winged Skimmer. State: Hagen (1861*, 1863, 1874, 1875), Banks
(1892), Calvert (1893), Ris (1910), N&W (1955), NW&M (2000), Donnelly
(2004b).
Atkinson (WMC), Brantley
(UG), Burke (UG), Camden (UG), Charlton (Br14, UG, UN), Chattahoochee (UG),
Chattooga (GB4), Clinch (GB3, UG, WMC), Cobb (GB3), DeKalb (UG), Early (GB3),
Echols (GB3, WMC), Glynn (By31), Harris (UG), Lanier (UG, UN), Laurens (GB3),
Lee (Ro24, RB3), Lowndes (UG, UN), McIntosh (UG), Thomas (GB1), Treutlen (GB3),
Walker (GDS3), Ware (RB, UN), Wayne (By31), Wheeler (WMC).
Uncommon
below the fall line and uncommon to rare above at ponds and temporary pools in
fields and forests. [May 8 - Aug 7].
Libellula
cyanea Fabricius, 1775. Spangled Skimmer. State: Muttkowski (1910*),
N&H (1929), Montgomery (1945), N&W (1955), NW&M (2000), Donnelly
(2004b).
Baldwin (SD, SP), Banks (UG),
Bartow (UG, UN), Bibb (UG), Burke (GB2), Carroll (GB2, WMC), Chattooga (GB2),
Clarke (UG), Clay (GB2), Cobb (GB, UG, UN), Coweta (GB2), Crawford / Bibb (SD),
Dade (GDS3), Dawson (GB, WMC, MW9), Early (GB2), Elbert (GB), Fannin (GB3),
Floyd (GB1), Franklin (UG, WMC), Fulton (UG), Gilmer (GB3), Gordon (SD), Greene
(MD3), Gwinnett (PA), Habersham (FSCA, GB1, JD, UN), Haralson (WMC), Harris
(GB), Henry (UG, UN), Houston (SD), Jasper (GB, GB2), Jones (WMC), Lamar (UG),
Lee (Ro24), Lumpkin (SD), Meriwether (GB1), Monroe (MD3), Morgan (UG), Murray
(GB3), Paulding (GB, GB1), Pickens (GB), Polk (GB, UG), Putnam (GB2), Rabun
(GBi, GB3), Rockdale (FS2, UN), Schley (GB3. RB3), Spalding (UG), Stephens
(UN), Talbot / Marion (SD), Taliaferro (GB), Taylor (SD), Union (SD), Walker
(SD, GDS3), Walton (UG), Washington (SD), White (SD, GB1, GB4, KT, WMC).
Common
above the fall line becoming uncommon below and absent from most of the eastern
and central coastal plain at ponds and marshes. [Apr 10 - Sep 2].
Libellula
flavida Rambur, 1842. Yellow-sided Skimmer. State: Hagen (1863*, 1874) and Banks (1892) as L.
plumbea, Calvert (1907), Ris (1910), N&W (1955), NW&M (2000),
Donnelly (2004b).
Bartow (GB1), Bibb (UG),
Bryan (UN), Burke (By31, GB3, UN, Wi34), Carroll (UN), Clarke (UG), Cobb (GB),
DeKalb (UG), Early (GB3), Emanuel (SD, GB4), Fannin (GB3, UN), Floyd (GB2),
Fulton (UG), Gordon (SD), Habersham (UN), Macon (SD), McDuffie (Wi34), Morgan
(MV3), Murray (KT), Oglethorpe (UG), Rabun (Da11, SD), Richmond (SD, UG),
Rockdale (FS2, UG), Schley (GB3), Sumter (RB), Taylor (SD), Union (SD).
Uncommon
throughout the state but mostly absent from the southernmost portion; in seeps
and very small streams. [Apr 21 - Oct 11].
Libellula
incesta Hagen, 1861. Slaty Skimmer. State: Hagen (1874*), Ris (1910), Byers (1927a),
N&W (1955), NW&M (2000), Donnelly (2004b).
Appling (GB, GB1), Baker
(UG), Baldwin (SP, UG), Bartow (GB, UG), Ben Hill (UG), Berrien (GB, GB3), Bibb
(UG), Brooks (GB1, UN), Bryan (UN), Bulloch (AH1), Burke (By31, Wi34), Butts
(MD3), Calhoun (UG), Catoosa (GB), Charlton (Br14, PA), Chatham (GB), Chattooga
(GB, GB2), Cherokee (GB), Clarke (JF, UG), Clay (GB2), Clayton (GB), Clinch
(UG), Cobb (GB), Coffee (RB), Colquitt (GB), Columbia (UN), Cook (MD3), Crawford
(UG), Crisp (GB2), Dade (GB2), Decatur (UG, UN), DeKalb (UG), Dooly (By31),
Douglas (GB), Early (GB2, SK4, UG), Elbert (GB), Emanuel (SD, GB1, GB4),
Fayette (KT), Floyd (GB), Forsyth (JF), Franklin (MD3), Gilmer (GB3), Glynn
(By31), Gordon (MD4), Grady (GB1), Greene (GB), Gwinnett (GB, PA), Habersham
(GBi, SD), Hall (MW9), Harris (GB, GB1), Henry (UG), Irwin (GBi, RB), Jasper
(GB), Jeff Davis (GB), Jenkins (JF, UG), Johnson (SD), Lanier (UG), Laurens
(By31, PA), Lee (Ro24, RB, UG), Lowndes (UN), Lumpkin (SD), Macon (GB, GB2),
McDuffie (Wi34), McIntosh (MV3), Meriwether (GB2), Miller (GB), Mitchell (UG),
Montgomery (GB1), Morgan (MD4), Murray (GB2), Paulding (GB), Pierce (JF4), Polk (GB), Rabun (GBi), Randolph (GB2), Richmond
(UG, UN), Rockdale (FS2, GB), Schley (RB3), Seminole (GB), Sumter (RB), Talbot
(SD), Taliaferro (JF4), Taylor (UG, UN), Telfair
(MD4), Tift (GBi, GB3), Toombs (GB1), Treutlen (GB1), Troup (GB), Twiggs (GB3),
Walker (GB2), Ware (RB), Wayne (By31), Wheeler (SD, WMC), White (SD, JF),
Whitfield (MD3), Wilcox (GB2), Wilkinson (SD).
Very
common statewide in puddles, ponds, lakes, and slower stretches of rivers and
streams. [May 6
- Nov 7].
Libellula
luctuosa Burmeister, 1839. Widow Skimmer. State: N&W (1955*), NW&M(2000), Donnelly
(2004b).
Baldwin (SP), Bartow (GB,
UG, UN), Burke (GB2), Butts (MD4), Catoosa (GB), Chattooga (GB, GB2, UN),
Clarke (UG), Clayton (GB), Cobb (GB), Dade (GB2), Dawson (GB), DeKalb (UG, UN),
Douglas (GB), Elbert (GB), Fannin (GB3), Floyd (GB, UG), Forsyth (GB, JF), Franklin
(UG, WMC), Fulton (UG, HS4), Gilmer (GB3), Glascock (MD3), Greene (GB),
Gwinnett (GB), Habersham (SD, UG), Hall (MW9), Harris (GB), Henry (UG, UN),
Houston (SD), Jasper (GB), Laurens (GB1), Lee (RB), Lumpkin (SD), Meriwether
(GB1), Morgan (UG), Newton (RT), Paulding (GB), Pickens (GB), Polk (GB), Putnam
(GB2), Rabun (GBi, SD, UG), Richmond (UG), Rockdale (FS2, GB), Stephens (UN),
Sumter (RB), Talbot (GB1), Taliaferro (JF4),
Union (JD),
Walker (SD, GDS3, GB), White (SD), Whitfield (MD4).
Very
common above the fall line, uncommon below in the western coastal plain and
absent from the central and eastern coastal plain, at marshes, ponds, and
lakes. [May 18
- Nov 2].
Libellula
needhami Westfall, 1943. Needham's Skimmer. State: N&W (1955), NW&M (2000), Donnelly
(2004b).
Camden (GB), Coffee (GB4),
Glynn (GB3, MW43*), Ware (UN).
Uncommon
and local along the immediate coast in freshwater marshes, salt marshes and
ponds. [Mar 22
- Oct 3].
Libellula
pulchella Drury, 1773. Twelve-spotted Skimmer. State: Hagen (1863, 1874, 1875), Calvert
(1893), Ris (1910), N&W (1955), NW&M (2000), Donnelly (2004b).
Baker (GB2), Baldwin (SP),
Carroll (GB), Chatham (Bu39*, Ca98), Chattooga (GDS3), Cherokee (GB), Clarke
(UG), Cobb (GB), Crawford / Bibb (SD), Dade (GB2), DeKalb (UN), Douglas (GB),
Fannin (JB3), Floyd (GB), Lamar (GB), Laurens (GB), Lee (Ro24), McDuffie
(Wi34), Rabun (SD), Richmond (AH2), Rockdale (GB), Walker (GDS3), Walton (UG).
Uncommon
to fairly common above the fall line, uncommon below in the western coastal plain
and absent otherwise, in marshes, ponds, and lakes. [Apr 24 - Oct 22].
Libellula
semifasciata Burmeister, 1839. Painted Skimmer. State: Hagen (1863, 1874, 1875), Ris (1910),
NW&M (2000), Donnelly (2004b).
Appling (UG), Baldwin (SP),
Bibb (UG), Brantley (JF4), Bryan (UN), Calhoun (MV3),
Carroll (GB2), Charlton (OO), Chatham (Ha61*, Ca98 as L. bifasciata ),
Chattooga (GB4), Clinch (WMC), Cobb (GB1, UG), Coffee (GB3), DeKalb (UG), Early
(GB4, MD4), Echols (WMC), Emanuel (KT, UG), Evans (WMC), Floyd (MD4), Glynn
(By31), Gordon (SD), Habersham (UN), Laurens (MD4), Lee (Ro24), Long (WMC),
Morgan (MV3), Murray (SD, GB3, KT), Pierce (SD, UG), Pike (UG), Rabun (UN),
Rockdale (FS2), Schley (RB3), Talbot / Marion (SD), Tattnall (WMC), Taylor
(SD), Telfair (WMC), Toombs (WMC), Walker (GDS3), Ware (UG), Wayne (OO), White
(GB4).
Uncommon
statewide in marshes, grassy wet areas, and ponds, more common below the fall
line. [Mar 21 -
Jul 30].
Libellula
vibrans Fabricius, 1793. Great Blue Skimmer. State: Ris (1910*), N&W (1955), NW&M
(2000), Donnelly (2004b).
Baldwin (SP), Bibb (UG),
Brooks (GB1), Bryan (UN), Bulloch (AH1), Burke (By31, UN, Wi34), Butts (MD4),
Calhoun (UG), Camden (UG), Charlton (Br14, OO), Chatham (GB), Cherokee (GB2),
Clarke (UG), Clinch (GB, UG), Colquitt (GB), Crisp (GB2), Decatur (UG, UN),
DeKalb (UG), Dooly (UG, UN), Dougherty (GB2), Early (SK4, UG), Echols (WMC),
Effingham (UN), Emanuel (GB4), Evans (UN), Floyd (GB), Fulton (UG), Glynn
(By31, UG), Henry (UG), Houston (SD), Irwin (GBi, RB), Jackson (UN), Jefferson
(UG), Jeff Davis (MD4), Johnson (SD), Lanier (UG, UN), Laurens (By31), Lee
(Ro24), Liberty (UG), Long (GB), Lowndes (UN), McIntosh (GB, UG), Miller (GB),
Montgomery (GB2), Murray (GB2), Paulding (GB2), Peach (UG), Rockdale (FS2),
Seminole (GB), Sumter (RB, UG), Taliaferro (JF4),
Tattnall (GB1),
Terrell (UG), Thomas (UN), Twiggs (GB3), Walker (GDS3, SG3), Ware (RB), Wayne
(OO), White (KT), Wilcox (GB2).
Fairly
common statewide at ponds, forest pools, and slow streams. [Apr 8 - Sep 19].
Miathyria
marcella (Selys in Sagra, 1857). Hyacinth Glider. State: NW&M (2000*),
Donnelly (2004b).
Camden (GB), Chatham (GB),
Crisp (GB2), Lanier (FSCA, GB4), Laurens (GB), Lowndes (MD4), Seminole (GB),
Sumter (GB2).
Uncommon
in the coastal plain, but can be locally abundant; at lakes and ponds with
water hyacinth or swarming nearby. [Apr 21 - Nov 5].
Nannothemis
bella (Uhler, 1857). Elfin Skimmer. State: Hagen (1863*, 1874, 1875), Banks (1892),
Calvert (1893), Ris (1911), Muttkowski (1910), N&H(1929),
Byers (1930). N&W (1955), NW&M (2000), Donnelly (2004b).
Bulloch (UN), Columbia
(FSCA), Richmond (SD, UN), Talbot / Marion (SD), Taylor (SD, GB4, MD4), Taylor
/ Schley (SD).
Rare
and local in bogs and marshes in the coastal plain, so far mostly near the fall
line. [Mar 5 -
May 17].
Orthemis
ferruginea (Fabricius, 1775). Roseate Skimmer. State: NW&M (2000), Donnelly (2004b).
Appling (GB), Atkinson
(MD4), Baker (GB1, GB2), Berrien (GB), Bryan (DS), Bulloch (AH1), Charlton
(PA), Chatham (FSCA, GBi97*), Cobb (GB), Colquitt (GB), Crisp (GB2, GB4),
Dougherty (SK), Early (GB4, SK4), Floyd (MD3), Glynn (GB), Jeff Davis (MD4),
Laurens (GB), McIntosh (GB), Meriwether (MD3), Miller (GB), Montgomery (MD4),
Muscogee (GB), Peach (GB), Seminole (GB2), Stewart (GB), Sumter (RB), Tattnall
(GB1, GB3), Tift (MD4), Wilcox (GB2).
Common
in the coastal plain, rare above but expanding slowly; at marshes, ponds, and a
few at lakes.
[May 17 - Dec 7].
Pachydiplax
longipennis (Burmeister, 1839). Blue Dasher. State: Hagen (1861*, 1863, 1874, 1875), Ris
(1911), N& WMC(1955), NW&M (2000), Donnelly (2004b).
Appling (GB), Atkinson (RB,
WMC), Baker (MV3, UG, UN), Baldwin (SP), Bartow (GB, UG), Ben Hill (WMC),
Berrien (GB), Bibb (GB, UG), Brantley (BB, UG), Brooks (UG), Bryan (GB, UN),
Bulloch (AH2, UN), Burke (By31, Wi34), Butts (MD4), Calhoun (MV3), Camden (UG),
Catoosa (GB), Charlton (Br14, GB4, OO, PA, UG, UN), Chatham (GB, H, UG, WMC),
Chattahoochee (UG), Chattooga (GB, GB2), Cherokee (GB), Clarke (UG), Clay (GB2),
Clayton (GB), Clinch (SD, UG, WMC), Cobb (GB), Coffee (RB), Colquitt (GB), Cook
(UG, UN), Coweta (GB), Crawford (UG), Crisp (GB2), Dade (GB2), Dawson (GB),
Decatur (UG, UN), DeKalb (UG, UN), Dodge (GB1), Dooly (By31, UN), Dougherty
(GB2), Douglas (GB), Early (GB2, SK4, UG, UN), Echols (WMC), Effingham (GB2,
UN), Elbert (GB), Emanuel (GB1), Evans (GB), Fannin (JB3), Floyd (GB, UG),
Forsyth (GB, JF), Franklin (GB, WMC), Fulton (UG), Gilmer (GB3), Glynn (By31,
GB, UG, UN, WMC), Gordon (MD4), Grady (UG), Gwinnett (By31), Habersham (UN),
Hall (MW9), Haralson (GB2), Harris (GB1), Heard (GB2), Henry (UG, UN), Houston
(UG), Irwin (GB3, RB), Jasper (GB2), Jeff Davis (GB), Jenkins (JF), Johnson
(SD), Jones (WMC), Lamar (GB), Lanier (UG, UN), Laurens (GB, PA), Lee (Ro24,
RB, RB3, UG), Liberty (GB, UG, UN), Long (GB), Lowndes (MD4, UG, UN, WMC),
Lumpkin (SD), Macon (GB, GB2), McDuffie (JF4),
McIntosh (GB,
MV3, UG), Meriwether (GB1), Miller (GB), Mitchell (UG), Montgomery (GB1),
Morgan (MV3), Murray (UN), Newton (MV3), Paulding (GB), Pickens (GB), Pierce
(SD), Polk (GB), Putnam (GB2, UG), Richmond (UG, UN), Rockdale (FS2, GB),
Seminole (GB), Stephens (UN), Stewart (GB), Sumter (RB, UG), Tattnall (WMC),
Taylor (SD, RB3, UG), Telfair (MD4, WMC), Thomas (GB1, UG, UN), Tift (RB),
Toombs (WMC), Treutlen (MD4), Troup (GB, UG), Turner (GB3, WMC), Twiggs (GB3),
Upson (UG), Walker (SD, GDS3, GB, GB2), Walton (UG), Ware (MV3, RB, UG), Wayne
(OO, UN), Wheeler (WMC), White (SD, KT), Whitfield (GB2, H), Wilcox (GB2),
Wilkinson (GB3), Worth (WMC).
Very common throughout the state
at puddles, marshes, ponds and lakes.
[Mar 21 - Nov 11].
Pantala
flavescens (Fabricius, 1798). Wandering Glider. State: Hagen (1861*, 1863, 1874, 1875),
Calvert (1893), Ris (1913), N&W (1955), NW&M (2000), Donnelly (2004b).
Baldwin (SP), Bulloch (AH1),
Carroll (GB), Chattahoochee (UG), Clarke (UG), Clayton (AH2), Cobb (GB), Coffee
(GB4, RB), Colquitt (GB), Dade (GB2), Dodge (MD4), Early (SK4), Floyd (GB,
GB2), Fulton (UG), Glynn (UG, WMC), Lanier (MD4), Lee (Ro24), Macon (GB),
McDuffie (Wi34), Miller (GB), Montgomery (MD4), Peach (GB), Rockdale (FS2),
Stewart (GB), Sumter (MD4), Union (Wi34), Wilcox (GB2) .
Fairly
common statewide at any still water habitat including puddles, uncommon at
rivers. Often seen in parking lots, highway intersections and sports fields,
especially in the fall. [Apr 22 - Dec 29].
Pantala
hymenaea (Say, 1839). Spot-winged Glider. State: N&W (1955), NW&M (2000), Donnelly
(2004b).
Bulloch (AH1), Clarke (UG),
Cobb (GB, GB3, GWQ71), Early (GB4), Floyd (MD3), Fulton (GWQ71), Irwin (RB),
Lee (Ro24*), Madison (UG), McDuffie (Wi34), Taylor (SD), Union (Wi34), Walker
(GDS3).
Uncommon
statewide at puddles and shallow ponds, and rarely at rivers. [May 11-Oct 12].
Perithemis tenera (Say, 1839). Eastern Amberwing. Includes seminole; State: Hagen (1875) and as P. domitia, Calvert
(1906), Ris (1910), Byers (1930), N&W (1955), NW&M(2000) as P. t.
seminole, Donnelly (2004b).
Appling (GB, GB1), Baker (GB2), Baldwin (SP, UG),
Bartow (GB), Ben Hill (GB2, WMC), Berrien (GB, GB3), Bibb (UG), Brooks (GB1),
Bulloch (AH1, AH2), Burke (Ha63*, Ha74 as P. domitia, By31), Butts
(MD4), Carroll (GB2, WMC), Catoosa (GB), Chatham (GB, UG), Chattooga (GB, GB2),
Clarke (JF, UG), Clayton (GB), Clinch (UG), Cobb (GB), Colquitt (GB, MD4, UG), Cook (UN),
Crawford (UN), Crisp (GB2), Dade (GB2), Dawson (GB), Decatur (UG), DeKalb (UG),
Dodge (GB1), Dougherty (GB2), Douglas (GB), Early (GB2, SK4), Elbert (GB),
Emanuel (GB1), Fayette (KT), Floyd (GB), Forsyth (GB, JF), Franklin (UG, WMC),
Fulton (UG), Gilmer (GB3), Glynn (GB2, UG), Gordon (GB2), Grady (UG, UN),
Gwinnett (GBi, PA), Habersham (GBi, SD), Hall (MW9), Hancock (MD4), Harris
(GB), Henry (UG), Houston (UG), Irwin (SD, GB3), Jasper (GB2), Jeff Davis (GB),
Johnson (SD), Jones (WMC), Lamar (GB), Laurens (GB), Lee (RB, UG), Liberty
(GB), Lowndes (GB, UN), Lumpkin (GB1), McDuffie (Wi34), McIntosh (GB, MV3),
Meriwether (GB1), Miller (GB), Montgomery (GB2), Morgan (UG), Paulding (GB),
Peach (GB), Pickens (GB), Pierce (JF4), Polk (GB), Putnam (UG),
Rabun (GBi), Richmond (UN), Rockdale (FS2, GB), Seminole (GB), Stephens (GB2),
Sumter (RB, UG), Talbot / Upson (GB2), Taliaferro (GB2), Tattnall (UN), Telfair
(MD4), Thomas (UG), Tift (GB3, UG), Toombs (GB1), Treutlen (MD4), Troup (GB),
Twiggs (GB3), Walker (GDS3, GB2), Ware (MV3), Washington (MD4), Wayne (By31),
Whitfield (GB2), Wilcox (GB2).
Common
across the state at marshes, ponds, and lakes, and along slower stretches of
large streams and rivers. [Apr 6 - Nov 3].
Plathemis
lydia (Drury, 1770). Common Whitetail. (Libellula of some
authors). State: Hagen
(1861, 1863, 1874, 1875) and Ris (1910) also as Libellula trimaculata, N&W (1955), NW&M
(2000), Donnelly (2004b).
Appling (GB), Baker (UG),
Baldwin (SP), Banks (GB1), Bartow (GB2, UN), Berrien (GB), Bibb (GB), Bulloch
(AH2), Butts (MD3), Calhoun (MV3), Camden (UG), Carroll (UG), Catoosa (GB, UN),
Charlton (UG), Chatham (Bu39* as L. trimaculata, Ca98), Chattooga (GB),
Cherokee (GB), Clarke (UG), Clay (GB2), Clayton (GB), Clinch (UG), Cobb
(GB),Colquitt (GB, MV3), Cook (MV3, UG), Coweta (UG), Crawford (UG), Crisp
(GB2), Dade (GDS3, GB2, UG), Dawson (GB, WMC, MW9), Decatur (GBi), DeKalb (UG),
Dooly (GB2), Early (GB4, SK4), Echols (WMC), Effingham (UG), Elbert (GB),
Emanuel (MD4), Fannin (JB3), Fayette (KT),
Floyd (By31, GB), Forsyth (JF, RT), Franklin (WMC), Fulton (UG), Gilmer
(GB3, WMC), Glynn (UG), Gordon (SD), Grady (UG), Greene (GB), Gwinnett (By31,
JF, UG), Habersham (SD, UG), Hall (MD3), Haralson (GB2), Harris (GB, MV3),
Houston (UG), Jasper (GB), Jeff Davis (UG), Jefferson (UG), Jones (MV3,
WMC), Lamar (GB, UG), Lanier (UG),
Laurens (By31, GB), Lee (Ro24, RB, UG), Liberty (UG), Long (WMC), Lowndes (UG),
Lumpkin (SD, UG), Macon (GB), McDuffie (Wi34), McIntosh (MV3, UG), Meriwether
(MV3, UG), Miller (GB), Mitchell (UG), Monroe (UG), Montgomery (MD4), Morgan
(MV3), Murray (SD, SR, WMC), Newton (MV3), Oconee (UG), Paulding (GB), Peach
(GB), Pickens (GB), Pierce (SD), Pike (UG), Polk (GB), Pulaski (WMC), Putnam
(GB2, UG), Rabun (GBi, UN), Richmond (UG, UN), Rockdale (FS2), Stephens (UG), Stewart (GB), Sumter (RB,
UG), Talbot (GB1), Taliaferro (GB2, UG), Tattnall (WMC), Taylor (SD, UG),
Thomas (UG), Tift (GBi, UG), Toombs (WMC), Troup (UG), Twiggs (GB3), Union
(SD), Upson (SD), Walker (GDS3, GB2, SR), Walton (UG), White (SD), Whitfield
(MD4), Wilcox (GB2), Worth (UG),
Abundant
and conspicuous statewide at any standing water, found uncommonly at slow
streams. [Mar
17 - Nov 3].
Sympetrum
ambiguum (Rambur, 1842). Blue-faced Meadowhawk. State: Selys & Hagen (1850*), Hagen
(1861, 1863, 1874, 1875) and Banks (1892) as Diplax albifrons, Muttkowski (1910), Ris (1911). N&H (1929),
Montgomery (1945), N&W (1955), Carle (1993), NW&M (2000), Donnelly
(2004b).
Baldwin (SP), Barrow (UG), Burke
(GB2, Wi34; Ta67), Carroll (GB3), Chattooga (UG), Clarke (Ta67, UN), Cobb (GB),
DeKalb (UG), Dougherty (UG), Floyd (GB2, MD3), Forsyth (JF), Franklin (UG),
Fulton (UG), Gordon (GB2), Gwinnett (PA), Laurens (GB4), Lee (Ro24, Ta67),
Lowndes (Ta67), Meriwether (SD), Monroe (GB4), Paulding (GB), Rockdale (FS2,
GB), Toombs (UG), Wheeler (SD). Rambur Type from
Georgia.
Fairly
common above the fall line, and uncommon in the northern section of the coastal
plain, at marshes and ponds. [Apr 24 - Nov 14].
Sympetrum
corruptum (Hagen, 1861). (Tarnetrum of some authors). Variegated
Meadowhawk. State: Ris
(1911*), Tai (1967), Carle (1993), N&W (1955), NW&M (2000), Donnelly
(2004b).
Clarke (UG), Glynn (WMC).
This species is common in
the western United States but rare in Georgia as well as in other eastern
states. [No date information]
Sympetrum
rubicundulum (Say, 1839). Ruby Meadowhawk. State: Carle (1993), NW&M (2000), Donnelly
(2004b).
Rabun (SD83*), Towns (KT),
Union (SD), White (D8)
Rare
in the extreme northern part of the state at marshes and ponds. It is known only from four records. [Jun 7 - Aug 29].
Sympetrum
semicinctum (Say, 1839). Band-winged Meadowhawk. State: NW&M (2000), Donnelly (2004b).
Towns (GBi90*, KT), Walker
(GDS3).
Rare
in the extreme northern part of the state in shallow flowing marshy areas. It is known only from two records. [May 24 - Aug 7].
Sympetrum
vicinum (Hagen, 1861). Autumn Meadowhawk. State: Ris (1911*), N&W (1955), Carle (1993),
NW&M (2000), Donnelly (2004b).
Baldwin (SP), Burke (Wi34),
Carroll (GB), Catoosa (GB2), Clarke (UG), Cobb (GB2), DeKalb (UG), Fannin
(GB3), Floyd (GB2), Greene (GB), Heard (GB2), Lumpkin (SD), Murray (GB3),
Oconee (UG), Paulding (GB2), Rabun (SD, GB3), Troup (JF4), Union (SD), Walker (MD3), Washington (SD), White
(UG).
Fairly
common but scattered at marshes and ponds above the fall line, rare just below
the fall line.
[Jun 7 - Nov 28].
Tramea
carolina (Linnaeus, 1763). Carolina Saddlebags. State: Hagen (1861*, 1863, 1874, 1875), Ris
(1913), Cuyler (1989b), N&W (1955), NW&M (2000), Donnelly (2004b).
Appling (GB4, UG), Atkinson
(MD4), Baker (GB1, MV3, UG), Baldwin (SP), Bartow (UG), Brantley (BB), Bryan
(GB1, UN), Bulloch (AH1, AH3), Burke (MD4), Calhoun (MV3), Charlton (OO, PA,
UG, UN), Chatham (GB, UG), Chattooga (GB2), Clinch (UG, WMC), Coffee (GB3, UG),
Cook (UG, UN), Crawford (UG, UN), Crisp (GB2), Dade (GDS3, GB2), Decatur (UN),
DeKalb (GB, UG), Effingham (UN), Emanuel (KT), Fannin (GB3), Floyd (GB),
Forsyth (GB), Fulton (HS4), Glynn (By31, GB, UG), Gwinnett (GB, PA), Hancock
(MD4), Harris (MV3), Houston (SD), Irwin (SD), Jasper (GB2), Jeff Davis (UG),
Johnson (UG), Lanier (GB3, UG, UN), Laurens (GB), Lee (Ro24, RB, RB3), Liberty
(UG), Long (GB4, WMC), Lowndes (UG, UN, WMC), McIntosh (MV3), Meriwether (MV3),
Mitchell (UG, UN), Monroe (MV3), Morgan (MV3), Newton (MV3), Pierce (SD),
Putnam (GB2), Richmond (UN), Rockdale (FS2), Seminole (GB), Talbot (GB1),
Tattnall (GB1, WMC), Taylor (SD, UG), Tift (UG), Toombs (WMC), Walker (GB2),
Ware (MV3, UG, UN), Wayne (By31, OO), Wheeler (WMC), White (SD), Whitfield
(GB2).
Common
statewide at marshes, ponds and lakes or as coastal migrants. [Mar 11-Dec 7].
Tramea
lacerata Hagen, 1861. Black Saddlebags. State: N&W (1955*), NW&M (2000), Donnelly
(2004b).
Baker (MV3), Baldwin (SP),
Berrien (GB), Bulloch (AH1, A3), Carroll (GB2), Catoosa (GB), Chattooga (GB2),
Clarke (UG), Cobb (GB), Coffee (RB), Colquitt (MD4), Crisp (GBi), Dade (GB2),
DeKalb (UN), Early (GB2, UG), Floyd (GB), Franklin (WMC), Glynn (WMC), Greene
(GB), Jasper (GB2), Lamar (GB), Laurens (GB), Lee (RB), Lumpkin (GB1), Macon
(GB), McDuffie (Wi34), McIntosh (GB4, MV3), Miller (GB), Monroe (MV3), Morgan
(MV3), Newton (MV3), Paulding (GB1), Peach (GB), Polk (GB), Putnam (GB2), Rabun
(GBi), Rockdale (FS2), Seminole (GB),
Sumter (RB), Tift (MD4), Walker (GDS3).
Common
to fairly common statewide at marshes, ponds and lakes or as coastal migrants. [Mar-24 - Nov 15].
Tramea
onusta Hagen, 1861. Red Saddlebags. State: Ris (1916), N&W (1955), NW&M (2000),
Donnelly (2004b).
Charlton (Br14, UN)
There are 8 specimens at
NMNH that could not be located for verification. The counties listed are:
Appling, Fulton, Jeff Davis & Sumter. Rare
vagrant south of the mountains, known only from few records at marshes, ponds,
or lakes. [No date information]
Doubtful Records and Synonyms
Previously published Georgia records of the
following are removed because of one or more of the following reasons:
misidentification, synonymy, indefinite locality, no recent collections,
vagrants, and/or out of known range.
Calopteryx apicale
Burmeister 1839.
It was listed for Burke County (Williamson 1934) and Charlton County (Bradley
1914). Hagen (1889) speculated on its synonymy, and Johnson (1973b) concluded
that it as a synonym of C. dimidiata.
Calopteryx splendens
(Harris, 1776). It was listed for the state by Hagen (1875), who mentioned “a
male in the Zurich Museum labeled Georgia, Abbot, probably erroneous.” Hagen
(1889) “could not find specimen collected by Abbot from Georgia in the Escher
Zollikofer coll.” Kirby (1890) said
that it was a female C. angustipennis. C. splendens is a European species highly unlikely to
be found in Georgia.
Calopteryx tricolor Burmeister, 1839 ( = Hetaerina
tricolor). It
was listed for the state by Selys (1853*, 1854a), Hagen (1861, 1863, 1874,
1875), Banks (1892), Calvert (1893, 1906).
It was listed for Charlton County by Bradley, (1914; as H. tricolor).
Byers (1930) lists it as a synonym of Hetaerina
titia. We consider it a synonym of H. titia.
Calopteryx virginica
Selys 1853. It was
listed for the state by Hagen (1861*, 1863). Hagen (1874) stated that it is a
synonym of C. aequabilis. This
was probably C. dimidiata,
since the nearest records for aequabilis
are Ohio and Pennsylvania (Westfall & May (2000)).
Hetaerina limbata
Selys 1853. This
was listed for the state by Muttowski (1910). Byers (1930) considered H.
limbata a subspecies of H. titia. Garrison (1990)
referred to it as a “race”. It is here considered synonymous with H.
titia.
Hetaerina
septentrionalis Selys, 1853. It was listed for the state by Selys (1853*,
1854a), Hagen (1861, 1863, 1874, 1875), (1892) and Muttkowski (1910). Kirby
(1890) synonymized it with H. titia.
Sylphus elegans
Hagen in Selys 1853. (= Calopteryx elegans). It was listed for the state
by Selys (1854a*, 1859, 1879a). Selys (1879a) further states that the female
type specimen is probably a synonym of Calopteryx angustipennis.
Lestes forcipatus
Rambur, 1842. It was
listed for the state by Selys (1862*), Hagen (1863), Banks (1892), Muttkowski
(1910). N&H (1929), Byers (1931). Bryan (UN), and Glynn (UN), and for Lee
County (Root 1924). T.W. Donnelly examined the 2 NMNH specimens and found them
to be L. australis (pers. comm.) He said he would not
"trust" the Root record from Lee County. He further stated, “Lee
County is in southwest Georgia. I would expect that this was L. australis
also.” Montgomery (1941) discussed
confusion between forcipatus and disjunctus in
Indiana. Walker (1952) said that the earlier records should be Lestes d.
australis. W&M (1996) listed it as “GA?” based on the senior
author’s preliminary list. It has been found near northern Georgia (see also
under expected species); but until a valid record is found, it is removed.
Lestes hamatus Hagen,
1861. It was listed for the state by Hagen (1874).
This is a synonym of Lestes forcipatus. See discussion under that
species.
Lestes uncatus Kirby 1890 (= Lestes
dryas Kirby 1890). It was listed for
the state by Calvert (1893). Cowley (1935) designated L. dryas as
the senior name to L. uncatus. L. dryas is listed in W&M (1996) as “GA?”
based on preliminary data provided by the senior author. It is now designated
as doubtful.
Agrion fontium Hagen, 1861. It was listed for
the state by Hagen (1863*). Hagen (1874) synonymized it with Argia
tibialis.
Aeshna abboti Hagen 1874. It was listed for the
state by Hagen (1874*). Hagen (1875) synonymized it with Corpyaeshna
ingens.
Aeshna quadriguttata Burmeister, 1839. It was listed
for the state by Hagen (1863*, 1874). Hagen (1875) synonymized it with Boyeria
vinosa.
Aeshna virens
Rambur 1842 (= Coryphaeshna
viriditas) Calvert, 1952. It was listed for the state by
Hagen (1874, 1875), Banks 1892, Calvert (1903b, 1906), Kirby (1890), Martin
(1908), Muttkowski (1910), and N&H (1929). Hagen (1874) implies that this
record is unverified, but states, “their occurrence in Georgia would not be
exceptional.” Hagen (1875) doubts the earlier records. Needham & Heywood
(1929) echoed the original record and list it as “Georgia?” It may have been a
vagrant, but since there are no records within the last century, we remove it.
Anax concolor Brauer 1865. This is listed in Muttkowski (1910)
explicitly for “Ga.”. Muttkowski also
lists longipes as “Mass. & Ohio to Fla.”, strongly implying
that both taxa occur in Georgia. This
citation is mysterious; Martin (1908) considered concolor only a
race of longipes, whereas Muttkowski, written immediately afterwards,
elevated it to a species without explanation. Modern workers
consider that North American citations of concolor refer to longipes,
possibly in a spotted juvenile coloration.
NW&M (2000) considers it to be either a subspecies or synonym of Anax
longipes. The status of true tropical concolor is
contentious. Garrison (1991-2004) considers it a separate species, but the concolor
form is tropical. We consider that
Georgia records of concolor should be attributed to longipes.
Cordulegaster diastatops (Selys 1854). This was listed for the state by N&W
(1955). When Carle (1983) described C. bilineata, he placed all
of the Georgia records into that species. According to Donnelly (2004a), the
nearest records of diastatops are from the highlands of
West Virginia.
Gomphus pilipes
Hagen in Selys 1858. This was listed
for the state Selys (1858*), Hagen (1861, 1863, 1874, 1875). This has
been synonymized with Arigomphus pallidus (Muttkowski, 1910).
Gomphus (Gomphurus)
crassus Hagen in Selys, 1878. This was listed for the state by Louton (1982) and NW&M (2000).
Louton (1982) shows a dot in northwestern Georgia, but there is no reference to
it in his text. This may have been an error. Donnelly (2004a) does not show it
in Georgia. Even though it is likely to occur in Georgia (see “Expected
Species”), it is removed from the list until a voucher specimen is secured.
Gomphus septima
Westfall 1956. This was listed for the
state by NW&M (2000). The source of the NW&M data is uncertain. The
senior author edited the distribution list in that publication and it was not
included on this list. Bick (2003) indicated that it was an error. It may occur
in Georgia, but no voucher specimen could be located. See also under “Expected
Species.”
Progomphus borealis
McLachlan in Selys, 1873.
This was listed for the state by Hagen (1874, 1875). Selys (1878a) states that
this record is that of P. obscurus.
Cordulia lateralis Burmeister, 1839. This was listed for the state by Hagen (1863*, 1874). Hagen (1875)
synonymized it with Epitheca cynosura.
Epitheca calverti Muttkowski 1915. Muttkowski (1915) described it from a
specimen collected at Thomasville, Georgia. According to Davis (1933), the
specimen was destroyed, but he thought it was E. semiaquea.
Epitheca spinigera
(Selys, 1871).
This was listed for the state by Selys (1874*), Banks (1892), Martin (1906),
Muttkowski (1910), and N&H (1929). The source of the original record is
unclear, but according to Donnelly (2004b), the nearest records are southern
Illinois, northern Ohio, Pennsylvania and New Jersey. It is removed from the
list.
Somatochlora semicircularis (Selys 1871). Martin (1906) listed it for the state, but
subsequently there have been no references to this record, which was probably
an error.
Celithemis leonora Westfall, 1952. This was
listed for the state by N&W (1955*), NW&M (2000), and Donnelly (2004b). It is removed due to synonymy with C. bertha
(Garrison 1991, 2004).
Diplax albrifrons
Charpentier, 1840. This
was listed for the state by Selys & Hagen (1850*), Hagen (1861, 1863, 1874,
1875) and Banks (1892). Ris (1911) synonymized it with Sympetrum ambiguum.
Erythemis
haematogastra (Burmeister 1839). This was listed for the state by Hagen (1861*,
1863,1874). Hagen (1874) quotes a communication from Abbot: “. . . saw a male in the coll of Mr. Escher
Zollikofer in the Zurich Museum.”
Hagen (1875) stated: “one male by Abbot; locality still doubtful.” Kirby
(1890), Muttkowski (1910) and Ris (1910), repeated Hagen’s statement. Kennedy (1923)
stated in a footnote that Williamson, in a personal communication, doubts the
Georgia record. NW&M (2000) mentioned the original Hagen record and stated
“it has not been reported subsequently from the state of Georgia.” The species
ranges from Cuba, Jamaica, and Mexico south to Paraguay. We remove it from the
list.
Erythrodiplax umbrata (Linnaeus, 1758). Band-winged Dragonet. This was listed for the state by
Hagen (1861*, 1863, 1868, 1875), Banks (1892), Calvert (1906), Muttkowski
(1910), Ris (1911), N&H (1929), Byers (1930), Borror (1942), N&W
(1955), NW&M (2000), and Donnelly (2004b). All of these refer to one male
of unknown locality recorded by Abbot in the early 19th century. Dunkle (1989)
reports no records north of Highlands County in Florida.
Ladona exusta (Say 1839). This was listed for the state by Calvert (1893*), Ris
(1910), N&W (1955). Prior to Bennefield (1965) coloration was the
primary means of identifying the species of Ladona, thus some
misidentifications probably occurred. This species is northern in range (see
range maps of Bennefield, 1965; Dunkle (2000), and Donnelly (2004b)) A Georgia
record would be unlikely. This record probably refers to L. deplanata.
Libellula bifasciata
Fabricius 1772.
This was mistakenly synonymized with Libellula
pulchella by
Calvert (1898). According to NW&M (2000) it is really a synonym of L
. semifasciata. Garrison (1991-2004) also comments about this.
Libellula camilla
Rambur 1842. This
was synonymized with Celithemis
eponina (Hagen, 1861).
Libellula jesseana
Williamson 1922.
Westfall (1943) reports specimens at Emory University Collection. The senior
author after many emails and phone calls could not locate them. There were no
more references to these specimens even by Needham & Westfall (1955). There
is a likelihood that L. jesseana is in Georgia (see expected
species), but until a voucher specimen is found it is removed from the list.
Libellula plumbea
Uhler, 1857. This
was listed for the state by Hagen (1863*, 1874) and Banks (1892). Byers (1930)
listed it as a synonym of Libellula
flavida.
Libellula trimaculata
DeGeer, 1773. This was
listed for the state by Hagen (1861, 1863, 1874, 1875), and Ris (1910), and for
Chatham County (Burmeister 1939*). Byers (1930) listed it as a synonym of Plathemis
lydia.
Nannophya maculosa Hagen, 1861 (= Erythrodiplax maculosa). This was listed
for the state by Hagen (1861*, 1863, 1874, 1875), Brauer (1868), Karsch (1889),
Kirby (1890), Banks (1892) as Nannothemis maculosa, and by
Muttkowski (1910), Ris (1911). This is a South American species. It resembles Nannothemis
bella, which is probably what the Georgia specimen was. Borror (1942)
states, “The source locality ‘Georgia’ given by Hagen et al. is incorrect.”
Perithemis domitia
Drury 1773 and P. seminole Calvert, 1907. These taxa are discussed
together since along with P. tenera;
their names have been used interchangeably since their introduction. This was
listed for the state by Hagen (1875), and for Burke County (Hagen 1863*, 1874).
Prior to Calvert’s (1906) description of P. domitia seminole,
many of the Perithemis records from Georgia were named P.
domitia. Ris (1911) raised P. seminole to specific rank.
Byers (1930) could not find any difference between the larva of P.
domitia and that of P. seminole. N&W (1955) listed P.
seminole and P. tenera as distinct, but commented that no
difference could be found between the larvae (based on reared specimens).
NW&M (2000) listed it as a subspecies of P. tenera. We
incorporate all of these references into P. tenera.
This category is presented here to assist future
researchers with locating new records. These species have a reasonable
probability of occurring in Georgia, based on verified records from Alabama,
Florida, North Carolina, South Carolina, or Tennessee, within 100 miles of the
Georgia border. Some of the species on this list are also included in the
doubtful records list, due to lack of a verified record.
Lestes congener Hagen 1861. Spotted Spreadwing. Tennessen, Harper & Krotzer (1995) report it from Madison
County in north central Alabama, and Marion and Tuscaloosa counties in west
central Alabama. They also report that the habitat is usually small ponds,
especially those formed from surface mining. Donnelly (2004c) shows dots in
northeastern Alabama, as well as eastern Tennessee. All of these records are on
the western side of the mountain divide. The dates in Alabama ranged from May
10-Nov 6. We expect it could be found in extreme northwestern Georgia.
Lestes forcipatus Rambur 1842. Sweetflag
Spreadwing. This species is discussed under “Doubtful
Records”. Currently there are no valid records from Georgia. Bick (1983a)
reported a specimen in the FSCA from Madison County, Alabama, which is about 60
miles west of Northwest Georgia. Tennessen, Harper and Krotzer (1995) listed it
also from Madison, coll. Aug 24. Byers (1931) reported it from Macon, North
Carolina. Walker (1952) reported it from Nashville and Cumberland Counties
Tennessee. Look for it across northern Georgia since the southern border of its
range according to Donnelly (2003c) extends along an east-west plane just north
of the Georgia border into North Carolina.
Enallagma sulcatum Williamson 1922. Golden Bluet. Bick (1983a) reported a specimen
in the FSCA from Covington County, Alabama. This is about 90 miles west of
southwest Georgia. Dunkle (1989) reported it from Florida to Mississippi.
Dunkle (1992) listed scattered records from Leon to Clay Counties in northern
Florida. Williamson (1922a), reported it also from South Carolina. Tennessen,
Harper and Krotzer (1995) reported the habitat as sand-bottomed lakes and ponds
with emergent vegetation along the edges with a flight season of June 1 to
August 31. If one can find the right habitat in southern Georgia, E.
sulcatum could be there.
Nehalennia
irene (Hagen, 1861). Sedge Sprite. White, Tennessen, Fox &
Carlson (1983) reported it from Greenville and Pickens Counties in northwestern
South Carolina. These records are a few hundred miles south of the contiguous
range of this species but it may occur in northeastern Georgia.
Aeshna constricta Say 1839. Lance-tipped Darner. Brimley (1908) reported it from Highlands, North Carolina, which is less
than 10 miles from the northeast corner of Georgia. This location is about 100
miles south of the contiguous range of this species but it could occur in
extreme northern Georgia.
Aeshna tuberculifera Walker 1908. Black-tipped Darner. Another common northeastern species with two western North Carolina
records displayed by (Donnelly (2004a). This one and the next one might turn up
in extreme northern Georgia.
Aeshna verticalis Hagen 1861. Green-striped Darner. Like the previous one, this common northern species has two western
North Carolina records (Donnelly, 2004a).
Arigomphus
submedianius (Williamson, 1914). Jade Clubtail. Tennessen, Harper and Krotzer (1995) listed
it from Jackson Co AL, coll. July 7. This county is adjacent to the northwest
corner of Georgia. This species has a central United States range, but might
show up in northwest Georgia.
Gomphus (Gomphurus) fraternus (Say, 1839). Midland Clubtail.
Donnelly (2004a) displayed dot-map records across southern Tennessee just north
of Alabama and also in south central North Carolina. It may show up in northern
Georgia.
Gomphus (Gomphurus) crassus Hagen in Selys, 1878. Handsome Clubtail.
Tennessen, Harper and Krotzer (1995) listed it from Jackson County, Alabama,
about 30 miles from northwest Georgia. Tennessen & Louton (1984) listed it
from Bedford County and Coffee Counties, Tennessee, locations that are about 50
miles from the northwest tip of Georgia. Specimens were collected between early
May and early July. There is a likelihood of it occurring in northwestern
Georgia.
Gomphus (Gomphurus) septima Westfall, 1956. Septima’s Clubtail. Bick (1983b) reported it from Bibb and Tuscaloosa Counties in
Alabama, and Chatham County, North Carolina. In 2003 he reported it from a
number of localities across North Carolina. Krotzer (2003) reported a
collection date of May 5, so it probably has a mid-spring flight season.
Donnelly (2004a) displayed its known records, which would indicate that there
is a gap in its distribution that runs just above the fall line through
Georgia. Its habitat is medium to large rocky rivers.
Gomphus (Gomphus)
cavillaris Needham 1902. Sandhill Clubtail. Dunkle (1989, 1992) summarized its distribution,
which includes records across the northern part of Florida. Donnelly (2004a)
displayed its records which include extreme southern Alabama, all of northern
Florida with a number of records just south of the Georgia border and a few in
southeastern North Carolina. According to Tennessen, Harper and Krotzer (1995),
it has been collected in clean sand-bottomed lakes.
Gomphus (Gomphus) descriptus Banks 1896. Harpoon Clubtail. There
are two western North Carolina records , which
are the southernmost for this species. It could possibly be found in extreme
northern Georgia.
Gomphus (Gomphus)
hodgesi Needham 1950. Hodges’ Clubtail. Donnelly (2004a) showed its range based on
collections from southeast Louisiana eastward through the panhandle of Florida
to just east of the Tallahassee area. Look for it in extreme southwest Georgia
in clean sandy streams.
Gomphus (Gomphus)
sandrius Tennessen 1983. Tennessee Clubtail. Tennessen (1983) reported it from Shelbyville,
Bedford Co, Tennessee, which is about 50 miles northwest of Georgia. The
habitat is a shallow bedrock stream about 25 meters wide.
Gomphus (Hylogomphus) abbreviatus Hagen in Selys 1878. Spine-crowned
Clubtail. Montgomery (1940) reported it from
Oconee County, South Carolina at Clemson, April 17-28. This is very near the
northeast border of Georgia.
Gomphus (Hylogomphus) viridifrons Hine (1901). Green-faced Clubtail. Louton (1982) reported it from Tennessee just about 50 miles north
of the Georgia border along the eastern border of TN. Tennessen, Harper and
Krotzer (1995) found it south of Birmingham, Alabama between April 24 and May
2. According to Donnelly (2004a) all of these records are on the west slope of
the Appalachian ridge; look for it in extreme northwestern Georgia.
Ophiogomphus aspersus Morse 1895. Brook Snaketail. Donnelly (2004a) showed its southernmost record in southwest North Carolina.
It could turn up in northern Georgia.
Ophiogomphus howei Bromley 1924. Pygmy Snaketail. Tennessen (1993a) discussed a record from Monroe County,
Tennessee, Tellico River, which is about 25 miles north of the northwestern
Georgia border. He also reported it from Sparta in northern NC on the east side
of mountains. These are the southernmost records according to Donnelly (2004a),
but it could extend across the border in north Georgia.
Ophiogomphus incurvatus alleghaniensis Carle 1982. Alleghany Snaketail. Carle (1982)
reported it from Tuscaloosa, Alabama and Blount County, Tennessee. Based on
Donnelly (2004a) dot map data, it could occur on the west slope of the
mountains in northwest Georgia.
Progomphus alachuensis Byers 1940. Tawny Sanddragon. Dunkle (1989, 1992)
summarized its distribution in Florida with the nearest record being from Baker
County, which is adjacent to the southeast border of Georgia. The senior author
has collected it in Florida on sand-bottom lakes with emergent vegetation along
the shoreline.
Progomphus bellei Knopf & Tennessen 1980. Belle’s Sanddragon. Bick (1983b and 2003)
summarized its distribution in the panhandle and Tallahassee area of Florida.
One record is within 10 miles of the southwestern border of Georgia. In Florida
the senior author found it in small sandy streams as well as sandy lakes and
ponds.
Stylurus potulentus (Needham, 1942). Yellow-sided
Clubtail. Donnelly (2004a) showed the dot map
records extending along the coast from southwestern Mississippi to just west of
Tallahassee Florida. It could occur in extreme southwestern Georgia. The senior
author found it in shallow, medium sized, sand bottomed, tannic stained streams
with lots of overhanging vegetation along the shoreline.
Stylurus townesi (Gloyd,
1936). Townes’ Clubtail. Donnelly (2004a) summarized its distribution in dot map
fashion with records in North Carolina, South Carolina, the western panhandle
of Florida, Alabama and Mississippi. Gloyd (1936), Montgomery (1940) and Bick
(2003) reported the South Carolina record from Greenville on August 22. The
senior author collected it at the same locality described above for S. potulentus
in southeast Mississippi in late July. If one could find suitable habitat below
the fall line, then both these species might be found.
Didymops floridensis Davis,
1921. Florida Cruiser. Dunkle (1989, 1992) and Donnelly (2004b) summarized its
distribution across northern and central Florida, with some records less than
25 miles from the southern Georgia border. According to Dunkle (1989) the usual
habitat is sand-bottomed lakes margined with maiden cane and often bald
cypress.
Macromia illinoiensis illinoiensis Walsh 1895. Illinois River Cruiser. Although the dot map (Donnelly, 2004b) appears to show it in northern
Georgia, the raw data provided by Donnelly (pers. comm.) do not show any
records within Georgia, but it has been collected very close to the
northwestern Georgia border. Some workers do not delineate the two subspecies
of M. illinoiensis. It may have been collected in Georgia
already but not differentiated.
Neurocordulia yamaskanensis (Provancher, 1875). Stygian
Shadowdragon. Although the
range of this species only extends to about 100 miles to the north and west of the
northern Georgia border (Donnelly 2004b), there is a remote possibility of it
occurring in northern Georgia. Tennessen et al (1995) stated it inhabits large
rivers and reservoirs.
Somatochlora calverti Williamson &
Gloyd, 1933. Calvert’s Emerald. Donnelly (2004b) showed it in
scattered locations in Florida, especially in the Tallahassee area just south
of southwest Georgia, plus a single collection site in the Aiken South Carolina
area near the Georgia border at Augusta. It should be in Georgia below the fall
line between these two populations. Collection dates were late summer.
Somatochlora hineana Williamson 1931. Hine’s Emerald. This rare
mid-western species is listed as “Threatened or Endangered.” There is one disjunct
record in northeastern Alabama (Tennessen, 1994b) in Jackson County, Robinson
Spring, 6.75 mi. N of Scottsboro on July 11. It is unlikely but possible in
Georgia based on this Alabama record.
Crocothemis servilia
(Drury, 1770). Scarlet Skimmer. Sprandel (1996) reported this Asian species from Gadsden
County, Florida, near Tallahassee, so it has spread northward since its first
discovery in south Florida in the mid 1970s. This is a very aggressive
competitor that can survive in less than perfect habitat.
Ladona julia
(Uhler, 1857). Chalk-fronted Corporal. Mark Cantrell (pers. comm.) reported it
from Jackson County, South Carolina, “only 11 miles from the Georgia border.”
He stated that the specimen was verified by Michael May in 2003.
Libellula jesseana Williamson 1932. Purple Skimmer. Dunkle (1989, 1992) summarized its distribution. Washington and
Clay Counties are the closest records to south-central Georgia. It inhabits
sand-bottom lakes with moderate vegetation along the shores. Westfall (1943)
reported specimens from Charlton and Macon counties, Georgia, at the Emory
University Museum, but did not report these records again in any future
publications. The specimens could not be located.
Macrodiplax balteata (Hagen, 1861). Marl Pennant. Byers (1934) reported it
from St. John’s County, Florida. Sprandel (1996) reported a Gadsden County,
Florida record. This species breeds in slightly brackish permanent pools along
the coast (NW&M, 2000) from the Barrier Island of
North Carolina along the coast all the way to Brownsville, Texas (Donnelly, 2004b).
In the west it inhabits marl ponds with high salinity. This type of habitat is
absent from Georgia but it could occur in the brackish areas along the Atlantic
coast.
Sympetrum internum Montgomery 1943. Cherry-faced Meadowhawk.
The single dot within North Carolina (Donnelly 2004b), based on raw data (pers.
comm.) is from Macon County, which is just north of the northeast corner of
Georgia.
Tramea calverti Muttkowski 1910. Striped Saddlebags.
This Neotropical species has been found throughout the
eastern United States probably as late summer vagrants (NW&M, 2000). There
are single records from northeastern Florida, coastal South Carolina and north
central North Carolina (Donnelly, 2004b). The South Carolina record was previously
reported as Tramea cophysa by Cuyler (1968), Needham &
Westfall (1955), White, Tennessen, Fox, & Carlson (1980) and NW&M
(2000). It may turn up in Georgia.
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